On the taxonomy of Philyra sagittifera (Alcock, 1896) and P. concinnus Ghani & Tirmizi, 1995 (Decapoda, Brachyura, Leucosiidae), with description of a new genus from the Indian Ocean Author Patel, Krupal Marine Biodiversity and Ecology Lab, Department of Zoology, The Maharaja Sayajirao University of Baroda, Vadodara, Gujarat, India. & Animal Taxonomy and Ecology Laboratory, Department of Life Sciences, Hemchandracharya North Gujarat University, Patan, Gujarat, India. Author Naderloo, Reza 0000-0002-5020-0962 Center of Excellence in Phylogeny of Living Organisms, School of Biology, University of Tehran, Tehran, Iran. rnaderloo @ ut. ac. ir; https: // orcid. org / 0000 - 0002 - 5020 - 0962 rnaderloo@ut.ac.ir Author Trivedi, Jigneshkumar Animal Taxonomy and Ecology Laboratory, Department of Life Sciences, Hemchandracharya North Gujarat University, Patan, Gujarat, India. Author Mitra, Santanu Zoological Survey of India, F. P. S. Building, 27 J. L. Nehru Road, Kolkata- 700016, West Bengal, India. text Zootaxa 2023 2023-08-16 5330 3 430 440 http://dx.doi.org/10.11646/zootaxa.5330.3.6 journal article 54886 10.11646/zootaxa.5330.3.6 3bd11a71-89b5-4e30-80c6-6078bb5be845 1175-5326 8254906 D8D32782-EE8C-4946-A873-F25F6F923FC6

TABLE 1. Morphological comparison between Tirmilyra n. gen. and its closely related genera (modified from Galil, 2009)

Characters	Tirmilyra n. gen.	Atlantophila Galil, 2009	Lyphira Galil, 2009	Pyrhila Galil, 2009	Hiplyra Galil, 2009
Carapace shape	Hexagonal	Suborbicular	Suborbicular	Suborbicular, globose	Suborbicular
Carapace regions	Distinct	Indistinct	Indistinct	Indistinct.	Indistinct
Dorsal surface	Three granulated or tuberculated ridges unite to from ‘arrow’ pointing forward	Glabrous	Glabrous	Variably granulate	Punctate and variably granulate
Hepatic facet	Broadly triangular tooth No triangular tooth projecting from lower projecting margin		No triangular tooth projecting	Lower margin prominently angular	No triangular tooth projecting.
Posterior margin	With pointed or blunt lateral tooth in male and female	Without lateral teeth	Without lateral teeth	With blunt lateral teeth in male	Without lateral teeth.
Chelipeds	Equal	Subequal	Subequal	Subequal	Subequal
Male pleon	Lacking subterminal denticle	Minute denticle subdistally	Denticle sub- terminally	Lacking subterminal denticle	Lacking subterminal denticle
G1	Shaft long, slender and straight	Shaft stout, sinuous and coiled on itself	Shaft long, dorsoventrally flattened	Shaft elongate, wide, dorsoventrally flattened.	Shaft wide, dorsoventrally flattened
Apical process	Long	Digitate, cornute	Short, cornute	Tubular, elongate	Minute