A New Species with Two New Subspecies of Rhinogobius (Teleostei: Gobiidae) from Yaeyama Group, the Ryukyu Islands, Japan Author Suzuki, Toshiyuki Osaka Museum of Natural History 1 - 23 Nagai Park, Higashi-Sumiyoshi-ku, Osaka 546 - 0034, Japan eviotatoshiyuki@gmail‛com Author Oseko, Naoharu Ginowan, Okinawa 901 - 2212, Japan Author Yamasaki, Yo Y. National Institute of Genetics 1111 Yata, Mishima, Shizuoka 411 - 8540, Japan Author Kimura, Seishi Graduate School of Bioresources, Mie University 1577 Kurimamachiya-cho Tsu, Mie 514 - 8507, Japan Author Shibukawa, Koichi Museum of Natural and Environmental History, Shizuoka 5762 Oya, Suruga-ku, Shizuoka, Shizuoka 422 - 8017, Japan text Bulletin of the Kanagawa Prefectural Museum (Natural Science) 2022 2022-03-29 51 9 34 journal article 296790 10.5281/zenodo.11267304 aef8aa96-4296-4a3d-a012-294e978cd29f 2189-6720 11267304 D4392EC9-CEE0-4414- AE64-296AAC207FEE

Table 3. Comparisons of 55 species-group taxa (species and subuspecies) of Rhinogobius with high or unknown vertebral counts

Species	Vertebrae	Longitudinal scales	Dark spot of D1	Bands or vertical rows Reference of dark spots on C.
Abbreviations: D1, first dorsal fin; C, caudal fin
R. albimaculatus	28	29–31	present	present	Chen et al., 1999a
R. aonumai aonumai	26–28	32–37	absent	present	This study
R. a. ishigakiensis	26–28	33–38	absent	present	This study
R. boa	27	30–33	present	present	Chen & Kottelat, 2005
R. bucculentus	unknown	40–44	absent	absent	Herre, 1927
R. carpenteri	28–29	34–40	absent	absent	Endruweit, 2017; Seale, 1910
R. changtinensis	27	28–30	present	present	Huang & Chen, 2007
R. cheni	27	34–36	present	present	Chen et al., 2008; Koumans, 1940; Nichols, 1931
R. chiengmaiensis	28	26–30	present	present	Chen et al., 2008; Fowler, 1934; Koumans, 1940
R. cliffordpopei	unknown	28–29	present	present	Nichols, 1925; Wu & Chen, 2008
R. coccinella	28	27–30	present	present	Endruweit, 2018
R. davidi	28	30–31	present	absent in male; present in female	Chen & Miller, 1998; Sauvage & Dabry de Thiersant, 1874
R. duospilus	27	29–31	present	present	Herre, 1935; Huang & Chen, 2007
R. filamentosus	27	30–33	present	present	Chen et al., 2008; Wu, 1939
R. flumineus	27–28	30–36	absent	absent or present	Akihito et al., 2013 & Errata; Mizuno, 1960b; This study
R. fukushimai	unknown	30–31	absent	present	Mori, 1934; Wu & Chen, 2008
R. genanematus	27	27–29	absent	present	Chen & Fang, 2006; Zhong & Tzeng, 1998
R. henryi	28	30–32	present	present	Chen et al., 2008; Herre, 1938
R. honghensis	28	32–34	present	present	Chen et al., 1999c
R. houheensis	30	37–40	present	absent	Wanghe et al., 2020
R. imfasciocaudatus	27–28	30–31	present	absent	Endruweit, 2018; Nguyen & Vo, 2005
R. immaculatus	27–28	29–31	absent	present	Li et al., 2018
R. lentiginis	27	30–32	present	absent in male; present in female	Chen & Miller, 1998; Wu & Zheng, 1985; Wu & Chen, 2008
R. lindbergi	27–28	30–32	present	present	Berg,1933; Sakai et al., 2000
R. lineatus	28	29–32	present	present	Chen et al., 1999a
R. linshuiensis	27–28	27–31	present	present	Chen & Miller, 2014; Chen et al., 2002
R. liui	29	35–39	present	present	Chen & Wu, 2008
R. longyanensis	27	30–32	present	absent	Chen et al., 2008
R. lungwoensis	28	31–34	present	present	Huang & Chen, 2007
R. maculagenys	27	32–34	present	present	Wu et al., 2018
R. maculicervix	28	29–32	present	present	Chen & Kottelat, 2000
R. maxillivirgatus	27	28–30	present	present	Xia et al., 2018
R. mekongianus	28	29–31	present	present	Chen et al., 1999a; Pellegrin & Fang 1940
R. milleri	27	28–31	present	present	Chen & Kottelat, 2003
R. multimaculatus	29	33–37	present	present	Chen et al., 1999b; Wu & Zheng, 1985
R. nammaensis	28	30–32	present	present	Chen & Kottelat, 2003
R. nanophyllum	28	29–32	present	present	Endruweit, 2018
R. ngutinhoceps	29	30–32	present	present	Endruweit, 2018
R. niger	27–28	35–37	present	absent in male; present in female	Huang et al., 2016
R. parvus	27	28–30	absent	absent in male; present in female	Huang et al., 2016; Luo, 1989
R. philippinus	unknown	36–40	absent	absent	Herre, 1927
R. phuongae	28–29	28–32	present	absent in male; present in female	Endruweit, 2018
R. ponkouensis	28	32–34	present	present	Huang & Chen, 2007
R. rubromaculatus	27–28	29–32	present	present	Lee & Chang, 1996; Wu & Chen, 2008
R. shennongensis	unknown	31–33	absent	present	Wu & Chen, 2008; Yang & Xie, 1983
R. sowerbyi	unknown	35–36	absent	absent	Ginsburg, 1917
R. sulcatus	27–28	26–29	present	present	Chen & Kottelat, 2005
R. szechuanensis	27	30–34	absent	present	Chen et al., 2008; Tchang, 1939; Wu & Chen, 2008
R. taenigena	27	29	present	present	Chen et al., 1999a
R. vermiculatus	28	32–34	present	present	Chen & Kottelat, 2003
R. wangchuangensis	27	28–30	present	absent in male; present in female	Chen & Miller, 2014; Chen et al., 2002
R. wangi	27	26–27	present	present	Chen & Fang, 2006
R. wuyanlingensis	27	30–32	present	present	Yang et al., 2008
R. xianshuiensis	27	29–32	present	present	Chen et al., 1999b; Wu & Chen, 2008
R. yaoshanensis	28	29–31	present	present	Chen et al., 2008; Luo 1989