| Bothynus ascanius (Kirby, 1819) Scarabaeus ascanius Kirby 1819: 399 | Brazil m | Endrödi 1985 |
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| Bothynus complexus Dechambre, 2006 Bothynus complexus Dechambre 2006: 229 | Brazil m | Dechambre 2006 |
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| Bothynus complanus (Burmeister, 1847) Podalgus complanus Burmeister 1847: 123 Bothynus simplicitarsis Bates 1888: 319 Bothynus monstrosus Bates 1888: 320 | Mexico, Guatemala, m, f Honduras, Panama, Colombia, Venezuela | Endrödi 1985, Ratcliffe 2003, Ratcliffe & Cave 2006, López-García et al. 2015 |
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| Bothynus cribrarius (Fairmaire, 1878) Scaptophilus cribrarius Fairmaire 1878: 266 | Brazil F | Endrödi 1985 |
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| Bothynus cunctator (Mannerheim, 1829) Scarabaeus cunctator Mannerheim 1829: 48 | Brazil, Bolivia, Paraguay m, f | Endrödi 1985 |
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| Bothynus cyclops (Burmeister, 1847) Corynoscelis cyclops Burmeister 1847: 127 | Guyana, Brazil, Paraguay m, f | Endrödi 1985 |
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| Bothynus cylindricus Arrow, 1937 Bothynus cylindricus Arrow 1937: 45 | Brazil, Argentina m | Endrödi 1985 |
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| Bothynus dasypleurus (Germar, 1824) Geotrupes dasypleurus Germar 1824: 116 | Brazil, Paraguay m | Endrödi 1985 |
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| Bothynus deiphobus (Burmeister, 1847) Podalgus deiphobus Burmeister 1847: 123 | Brazil m, f | Endrödi 1985 |
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| Bothynus entellus (LePeletier & Serville, 1828) Scarabaeus entellus LePeletier & Serville, 1828: 347 Scarabaeus glaucon Perty 1830: 45 Scarabaeus glycon Perty 1830: 214 Strategus montesuma Hope 1837: 88 | Peru, Brazil m, f | Endrödi 1985 |
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| Bothynus exaratus (Burmeister, 1847) Podalgus exaratus Burmeister 1847: 124 Scaptophilus exaratus argentinus Ohaus 1910a: 183 | Brazil, Paraguay, Argentina m, f | Endrödi 1985 |
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| Bothynus herbivorus (Arrow, 1937) Ligyrus herbivorus Arrow 1937: 44 Bothynus perforatus Arrow 1937: 45 | Colombia, Venezuela m, f | Endrödi 1985, López- García et al. 2015 |
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| Bothynus herteli Endrödi, 1968 Bothynus herteli Endrödi 1968: 173 | Guyana m, f | Endrödi 1985 |
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| Bothynus horridus Endrödi, 1968 Bothynus horridus Endrödi 1968: 171 | Brazil, Paraguay, Argentina m, f | Endrödi 1985, Abadie et al. 2008 |
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| Bothynus laevipennis Arrow, 1937 Bothynus laevipennis Arrow 1937: 46 | Brazil, Bolivia, Argentina m, f | Endrödi 1985 |
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| Bothynus lancifer Dechambre, 1981 Bothynus lancifer Dechambre 1981: 126 | Peru m | Dechambre 1981 |
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| Bothynus laticifex Burmeister, 1847 Bothynus laticifex Burmeister 1847: 115 | Brazil, Argentina m, f | Endrödi 1985, Abadie et al. 2008 |
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| Bothynus medon (Germar, 1824) Geotrupes medon Germar 1824: 115 Bothynus ascanius Burmeister 1847: 116 | Colombia, Brazil, Paraguay, m, f Bolivia, Argentina | Endrödi 1985, Abadie et al. 2008 |
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| Bothynus minor Steinheil, 1872 Bothynus minor Steinheil 1872: 561 | Argentina m, f | Endrödi 1985, Abadie et al. 2008 |
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| Bothynus niger (Blanchard, 1842) Cratocnemus niger Blanchard 1842: plate 11 | Argentina? | Blanchard 1842 |
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| Ligyrus bicorniculatus Casey 1915: 198 Ligyrus rubidus Casey 1915: 198 Ligyrus lucublandus Casey 1915: 199 Ligyrus curtipennis Casey 1915: 199 Ligyrus effetus Casey 1915: 200 Ligyrus farctus Casey 1915: 200 Ligyrus arizonensis Casey 1915: 201 Ligyrus brevipes Casey 1915: 202 Ligyrus laevicauda Casey 1915: 202 Ligyrus laticollis Casey 1915: 203 Ligyrus scitulus Casey 1915: 203 | | |
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| Tomarus gyas Erichson, 1848 Tomarus gyas Erichson 1848: 561 Ligyrus amazonicus Arrow 1914: 273 | Mexico, Guatemala, m, f Honduras, El Salvador, Nicaragua, Costa Rica, Panama, Suriname, Colombia, Venezuela, Ecuador, Peru, Bolivia, Brazil, Argentina | Endrödi 1985, Ratcliffe & Dechambre 1983, Ratcliffe & Cave 2006, López-García et al. 2015 |
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| Tomarus laevicollis (Bates, 1888) Ligyrus laevicollis Bates 1888: 316 Ligyrus bryanti Rivers 1891: 97 | Mexico, Guatemala, El m, f Salvador, Honduras, Nicaragua, Costa Rica | Endrödi 1985, Ratcliffe & Cave 2006, Ratcliffe et al. 2013 |
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| Tomarus maimon Erichson, 1847 Tomarus maimon Erichson 1847: 96 Heteronychus fossator Burmeister 1847: 101 | Colombia, Ecuador, Peru, m, f Brazil, Chile | Endrödi 1985, Ratcliffe & Dechambre 1983 |
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| Tomarus maternus (Prell, 1937) Ligyrus maternus Prell 1937b: 89 | Honduras, Nicaragua, Costa m, f Rica, Colombia, Venezuela, Ecuador, Peru, Bolivia | Endrödi 1985, Escalona & Joly 2006, Ratcliffe & Cave 2006 |
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| Tomarus nasutus (Burmeister, 1847) Podalgus nasutus Burmeister 1847: 120 Ligyrus pygidialis Bates 1888: 317 | Mexico, Guatemala, El m, f Salvador, Honduras, Nicaragua, Costa Rica, Panama, Jamaica | Endrödi 1985, Ratcliffe 2003, Ratcliffe et al. 2013 |
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| Tomarus neglectus (LeConte, 1847) Bothynus neglectus LeConte 1847: 87 | USA m, f | Endrödi 1985 |
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| Tomarus peruvianus (Endrödi, 1970) Ligyrus peruvianus Endrödi 1970: 106 | Peru, Ecuador m, f | Endrödi 1985 |
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| Tomarus pullus (Prell, 1937) Ligyrus pullus Prell 1937: 89 | Colombia, Venezuela m, f | Endrödi 1985, Escalona & Joly 2006, López- García et al. 2015 |
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| Tomarus pumilus (Prell, 1937) Ligyrus pumilus Prell 1937: 90 | Argentina m | Ratcliffe et al. 2013 |
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| Tomarus relictus (Say, 1825) Scarabaeus relictus Say 1825: 194 Ligyrodes clypealis Casey 1915: 181 Ligyrodes parviceps Casey 1915: 182 Ligyrodes quadripennis Casey 1915: 182 Ligyrodes relictus vernicicollis Casey 1915: 183 Ligyrodes dawsoni Casey 1915: 334 | Canada, USA m, f | Endrödi 1985, Bousquet et al. 2013 |
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| Tomarus rosettae (Endrödi, 1968) Ligyrus rosettae Endrödi 1968: 68 | Argentina m, f | Endrödi 1985 |
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| Tomarus rostratus Dupuis, 2014 Tomarus rostratus Dupuis 2014: 1 | Peru m | Dupuis 2014 |
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| Tomarus rubripes (Boheman, 1858) Podalgus rubripes Boheman 1858: 57 | Uruguay m | Endrödi 1985 |
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| Tomarus sallaei (Bates, 1888) Ligyrus sallaei Bates 1888: 318 Ligyrus propinquus Casey 1915: 183 Ligyroides aztecus Casey 1915: 183 | USA, Mexico, Guatemala, m, f El Salvador, Honduras, Nicaragua, Costa Rica | Endrödi 1985, Ratcliffe & Cave 2006, Ratcliffe et al. 2013 |
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| Tomarus selanderi (Cartwright, 1959) Bothynus selanderi Cartwright 1959: 537 | Mexico m, f | Endrödi 1985, Ratcliffe et al. 2013 |
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| Tomarus similis (Endrödi, 1968) Ligyrus similis Endrödi 1968: 166 | Panama, Venezuela, m, f Trinidad, Guyana, French Guiana, Colombia, Ecuador, Peru | Endrödi 1985, Ratcliffe 2003, López-García et al. 2015 |
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| Tomarus subtropicus (Blatchley, 1922) Ligyrus subtropicus Blatchley 1922: 20 Ligyrus blatchleyi Cartwright 1944: 34 | USA m, f | Endrödi 1985 |
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| Tomarus villosus (Burmeister, 1847) Podalgus villosus Burmeister 1847: 120 Ligyrus patagonus Steinheil 1872: 560 | Peru, Argentina, Uruguay, m, f Chile | Endrödi 1985 |
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| Acknowledgments | | |
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