| Bothynus nyx Ratcliffe, 2010 Bothynus nyx Ratcliffe 2010a: 105 | Bolivia | m | Ratcliffe 2010a |
|---|
| Bothynus quadridens (Taschenberg, 1870) Corynoscelis quadridens Taschenberg 1870: 185 | Costa Rica, Panama, Colombia, Venezuela, Ecuador | m, f | Endrödi 1985, Ratcliffe 2003 |
|---|
| Bothynus stenelus (Burmeister, 1847) Corynoscelis stenelus Burmeister 1847: 127 | Brazil | m, f | Endrödi 1985, Abadie et al. 2008 |
|---|
| Bothynus striatellus (Fairmaire, 1878) Scaptophilus striatellus Fairmaire 1878: 264 Scaptophilus fabius Fairmaire 1878: 264 | Brazil, Bolivia, Argentina | m, f | Endrödi 1985, Abadie et al. 2008 |
|---|
| Bothynus tricornis Arrow, 1937 Bothynus tricornis Arrow 1937: 44 | Brazil | m, f | Endrödi 1985, Abadie et al. 2008 |
|---|
| Bothynus validus (Burmeister, 1847) Podalgus validus Burmeister 1847: 125 | Brazil, Argentina | m, f | Endrödi 1985 |
|---|
| Bothynus villiersi Endrödi, 1968 Bothynus villiersi Endrödi 1968: 170 | Brazil, Paraguay | m, f | Endrödi 1985, Abadie et al. 2008 |
|---|
| Collagenus dasysternus Ratcliffe & Hardy, 2005 Collagenus dasysternus Ratcliffe & Hardy 2005: 144 | Venezuela | m | Ratcliffe & Hardy 2005 |
|---|
| Coscinocephalus cribrifrons (Schaeffer, 1906) Anoplocephalus cribrifrons Schaeffer 1906: 260 | USA, Mexico | m, f | Ratcliffe et al. 2013 |
|---|
| Coscinocephalus tepehuanus Morón & Ratcliffe, 1996 Coscinocephalus tepehuanus Morón & Ratcliffe 1996: 50 | USA, Mexico | m, f | Morón & Ratcliffe 1996, Ratcliffe et al. 2013 |
|---|
| Denhezia caucacola Dechambre, 2006 Denhezia caucacola Dechambre 2006: 227 | Colombia, Ecuador | m, f | Dechambre 2006 |
|---|
| Diloboderus abderus (Sturm, 1826) Scarabaeus abderus Sturm 1826: 66 Scarabaeus mentor Guérin-Meneville 1841: plate 23 | Peru, Brazil, Argentina, Uruguay | m, f | Endrödi 1985, Abadie et al. 2008, Ratcliffe et al. 2015 |
|---|
| Endroedianibe bozzolai Chalumeau, 1981 Endroedianibe bozzolai Chalumeau 1981: 221 | Dominican Republic, Haiti | m, f | Chalumeau 1981, Ratcliffe & Cave 2011 |
|---|
| Endroedianibe xerozoyphius Ratcliffe & Cave, 2011 Endroedianibe xerozoyphius Ratcliffe & Cave 2011: 4 | Dominican Republic | m | Ratcliffe & Cave 2011 |
|---|
| Eremobothynus bicuspis Endrödi, 1970 Eremobothynus bicuspis Endrödi 1970: 107 | Argentina | m, f | Endrödi 1970, Endrödi 1985 |
|---|
| Eremobothynus cornutus (Steinheil, 1872) Scaptophilus cornutus Steinheil 1872: 561 | Argentina | m, f | Endrödi 1985 |
|---|
| Eremobothynus hirsutus Grossi & Abadie, 2008 Eremobothynus hirsutus Grossi & Abadie 2008: 57 | Argentina | m | Grossi & Abadie 2008 |
|---|
| Euetheola humilis (Burmeister, 1847) Heteronychus humilis Burmeister 1847: 93 Ligyrus rugiceps LeConte 1856: 21 Dyscinetus parvus Casey 1915: 172 Dyscinetus hondurana Casey 1915: 188 | USA, Mexico, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Ecuador, Venezuela, Guyana, Brazil, Bolivia, Paraguay, Uruguay, Argentina | m, f | Endrödi 1985, Ratcliffe 2003, Ratcliffe & Cave 2006, Ratcliffe et al. 2013 |
|---|
| Euetheola latipennis Arrow, 1911 Euetheola latipennis Arrow 1911: 166 | Brazil, Bolivia, Paraguay, Argentina | m, f | Endrödi 1985 |
|---|
| Euetheola subglabra (Schaeffer, 1909) Ligyrus subglaber Schaeffer 1909: 383 | USA, Mexico | m, f | Endrödi 1985, Ratcliffe et al. 2013 |
|---|
| Gillaspytes janzeni Howden, 1980 Gillaspytes janzeni Howden 1980: 91 | Mexico | m, f | Howden 1980, García- Morales & Reyes-Castillo 2004 |
|---|
| Gorditus rotundus Ratcliffe, 2010 Gorditus rotundus Ratcliffe 2010b: 59 | Cuba | m | Ratcliffe 2010b |
|---|
| Heteroglobus obesus Dupuis & Dechambre, 2008 Heteroglobus obesus Dupuis & Dechambre 2008: 160 | Peru | m | Dupuis & Dechambre 2008 |
|---|
| Heteronychus arator Fabricius, 1775 Scarabaeus arator Fabricius 1775: 18 | Brazil | m, f | Endrödi 1985, Abadie et al. 2008 |
|---|
| Hiekeianus halapinus Ratcliffe, 2014 Hiekeianus halapinus Ratcliffe 2014: 668 | Peru | f | Ratcliffe 2014 |
|---|
| Hiekeianus tuberculatus Endrödi, 1978 Hiekeianus tuberculatus Endrödi 1978: 81 | Paraguay | m, f | Endrödi 1985, Abadie et al. 2008 |
|---|
| Hylobothynus arnaudi Dechambre, 1979 Hylobothynus arnaudi Dechambre 1979: 104 | Guyana, French Guiana, Surinam | m, f | Ratcliffe 1981b, Ratcliffe & Dechambre 1983, Ponchel 2011 |
|---|
| Hylobothynus columbianus Endrödi, 1969 Hylobothynus columbianus Endrödi 1969: 24 | Colombia, Ecuador, Peru | m | Endrödi 1985, Ratcliffe 1981b |
|---|
| Hylobothynus marielae Abadie, 2011 Hylobothynus marielae Abadie 2011: 159 | Brazil | m | Abadie 2011 |
|---|
| Hylobothynus obesus Ohaus, 1910 Hylobothynus obesus Ohaus 1910b: 678 | Colombia, Ecuador, Peru, Bolivia | m, f | Ratcliffe 1981b, Endrödi 1985, Abadie 2011 |
|---|
| Hylobothynus pontis Ratcliffe, 1981 Hylobothynus pontis Ratcliffe 1981b: 303 | Guyana, Brazil | m, f | Ratcliffe 1981b, Ponchel 2011 |
|---|
| Indieraligus michelii Dechambre, 1979 Indieraligus michelii Dechambre 1979: 102 | Puerto Rico | m | Ratcliffe & Dechambre 1983, Ratcliffe & Cave 2015 |
|---|
| Neoryctes galapagoensis (Waterhouse, 1845) Oryctes galapagoensis Waterhouse 1845: 26 | Ecuador | m | Endrödi 1985, Cook et al. 1995 |
|---|
| Neoryctes linelli Mutchler, 1938 Neoryctes linelli Mutchler 1938: 10 | Ecuador | m, f | Mutchler 1938, Endrödi 1985, Cook et al. 1995 |
|---|
| Neoryctes moreti Dechambre, 1991 Neoryctes moreti Dechambre 1991: 8 | Ecuador | m, f | Dechambre 1991, Cook et al. 1995 |
|---|
| Neoryctes similis Ratcliffe, 2014 Neoryctes similis Ratcliffe 2014: 670 | Ecuador | m, f | Ratcliffe 2014 |
|---|
| Neoryctes williamsi Cook, Howden, & Peck, 1995 Neoryctes williamsi Cook, Howden, & Peck 1995: 186 | Ecuador | m | Cook et al. 1995 |
|---|
| Orizabus amalgamatus Ratcliffe & Cave, 2010 Orizabus amalgamatus Ratcliffe & Cave 2010a: 2 | Mexico | m, f | Ratcliffe & Cave 2010a |
|---|
| Orizabus australis Ratcliffe & Solís, 2013 Orizabus australis Ratcliffe & Solís 2013: 2 | Costa Rica | m, f | Ratcliffe & Solís 2013 |
|---|
| Orizabus batesi Prell, 1914 Orizabus batesi Prell 1914: 203 | Mexico | m, f | Endrödi 1985 |
|---|
| Orizabus botox Ratcliffe & Cave, 2006 Orizabus botox Ratcliffe & Cave 2006: 203 | Honduras | m, f | Ratcliffe & Cave 2006 |
|---|
| Orizabus brevicollis Prell, 1914 Orizabus brevicollis Prell 1914: 204 | USA, Mexico | m, f | Endrödi 1985 |
|---|
| Orizabus clunalis (LeConte, 1856) Aphonus clunalis LeConte 1856: 23 Orizabus cultripes Fairmaire 1878: 261 Orizabus sallei Fairmaire 1878: 262 Orizabus marginatus Fairmaire 1878: 262 Orizabus snowii Horn 1885: 124 Orizabus ponderosus Casey 1915: 225 Orizabus fontinalis Casey 1915: 225 | USA, Mexico, Guatemala, El Salvador, Honduras | m, f | Endrödi 1985, Ratcliffe et al. 2013 |
|---|
| Orizabus cuernavacensis Delgado & Deloya, 1990 Orizabus cuernavacensis Delgado & Deloya 1990: 301 | Mexico | m, f | Delgado & Deloya 1990, Ratcliffe et al. 2013 |
|---|
| Orizabus dechambrei Morón, Tapia, & Aragón, 2003 Orizabus dechambrei Morón, Tapia, & Aragón 2003: 4 | Mexico | m, f | Morón et al. 2003, Ratcliffe et al. 2013 |
|---|
| Orizabus delgadoi Ratcliffe & Cave, 2010 Orizabus delgadoi Ratcliffe & Cave 2010a: 4 | Mexico | m | Ratcliffe & Cave 2010a, Ratcliffe et al. 2013 |
|---|
| Orizabus endrodianus Morón, 1981 Orizabus endrodianus Morón 1981b: 131 | Mexico | m, f | Morón 1981 |
|---|
| Orizabus epithecus Ratcliffe & Cave, 2010 Orizabus epithecus Ratcliffe & Cave 2010a: 5 | Mexico | m | Ratcliffe & Cave 2010a |
|---|
| Orizabus fairmairei (Bates, 1888) Cheiroplatys fairmaire Bates 1888: 321 | Mexico | m, f | Endrödi 1985, Ratcliffe et al. 2013 |
|---|
| Orizabus hondurensis Ratcliffe & Cave, 2006 Orizabus hondurensis Ratcliffe & Cave 2006: 208 | Honduras | m, f | Ratcliffe & Cave 2006 |
|---|
| Orizabus isodonoides Fairmaire, 1878 Orizabus isodonoides Fairmaire 1878: 263 | Mexico | m, f | Endrödi 1985, Ratcliffe et al. 2013 |
|---|
| Orizabus ligyroides Horn, 1885 Orizabus ligyroides Horn 1885: 125 | USA, Mexico | m, f | Ratcliffe et al. 2013 |
|---|
| Orizabus mcclevei Warner, 2011 Orizabus mcclevei Warner 2011: 11 | USA, Mexico | m, f | Warner 2011, Ratcliffe et al. 2013 |
|---|
| Orizabus mezclus Ratcliffe & Cave, 2010 Orizabus mezclus Ratcliffe & Cave 2010a: 6 | Guatemala | m | Ratcliffe & Cave 2010a |
|---|
| Orizabus pinalicus Warner, 2011 Orizabus pinalicus Warner 2011: 7 | USA | m, f | Warner 2011 |
|---|
| Orizabus puchicus Ratcliffe & Cave, 2006 Orizabus puchicus Ratcliffe & Cave 2006: 210 | Honduras | m, f | Ratcliffe & Cave 2006 |
|---|
| Oxyligyrus spitzi Endrödi, 1972 Oxyligyrus spitzi Endrödi 1972: 25 | Brazil, Paraguay | m | Endrödi 1985, Abadie et al. 2008 |
|---|
| Oxyligyrus rostratus (Burmeister, 1847) Chalepus rostratus Burmeister 1847: 82 | Guyana, Brazil | m, f | Endrödi 1985, Abadie et al. 2008 |
|---|
| Oxyligyrus zoilus (Oliver, 1789) Scarabaeus zoilus Oliver 1789: 45 Oxyligyrus punctulatus Endrödi 1966: 537 | Guyana, French Guiana, Colombia, Brazil, Bolivia, Paraguay | m, f | Endrödi 1985, Abadie et al. 2008, Ponchel 2011 |
|---|
| Parapucaya amazonica Prell, 1934 Parapucaya amazonica Prell 1934: 162 | Honduras, Panama, Colombia, Ecuador, Peru, Brazil | m, f | Endrödi 1969 |
|---|
| Parapucaya nodicollis (Kirsch, 1873) Cyclocephala nodicollis Kirsch 1873: 344 | Colombia, Ecuador, Peru | m, f | Endrödi 1985, Carvajal et al. 2011 |
|---|
| Pentodina peruviana Endrödi, 1968 Pentodina peruviana Endrödi 1968: 164 | Peru, Bolivia | f | Endrödi 1985 |
|---|
| Philoscaptus bonariensis (Burmeister, 1847) Podalgus bonariensis Burmeister 1847: 118 | Brazil, Uruguay, Paraguay, Argentina | m, f | Endrödi 1985, Abadie et al. 2008, Morón & Grossi 2015 |
|---|
| Piscoperus paracanicola Ratcliffe & Giraldo, 2014 Piscoperus paracanicola Ratcliffe & Giraldo 2014: 105 | Peru | m | Ratcliffe & Giraldo 2014 |
|---|
| Pucaya castanea Ohaus, 1910 Pucaya castanea Ohaus 1910b: 675 Pucaya columbiana Beck 1942: 47 | Panama, Colombia, Ecuador | m, f | Endrödi 1985, Restrepo- Giraldo et al. 2003, López-García et al. 2015 |
|---|
| Pucaya pulchra Arrow, 1911 Pucaya pulchra Arrow 1911: 167 Pucaya punctata Endrödi 1968: 165 | Colombia, Ecuador | m, f | Endrödi 1985, Carvajal et al. 2011, López-García et al. 2015 |
|---|
| Saccharoscaptus laminifer (Dechambre, 1979) Philoscaptus laminifer Dechambre 1979: 103 | Brazil, Bolivia | m, f | Dechambre 1979, Morón & Grossi 2015 |
|---|
| Thronistes rouxi Burmeister, 1847 Thronistes rouxi Burmeister 1847: 177 | Brazil, Uruguay, Argentina | m, f | Endrödi 1985, Abadie et al. 2008 |
|---|
| Tomarus adoceteus Ratcliffe & Cave, 2010 Tomarus adoceteus Ratcliffe & Cave 2010b: 7 | Bahamas, Cayman Islands | m, f | Ratcliffe & Cave 2010b, Ratcliffe & Cave 2015 |
|---|
| Tomarus bidentulus (Fairmaire, 1892) Ligyrus bidentulus Fairmaire 1892: 244 | Argentina | f | Endrödi 1985 |
|---|
| Tomarus bituberculatus (Palisot de Beauvois, 1811) Scarabaeus bituberculatus Palisot de Beauvois 1811: 103 Ligyrus latifovea Bates 1888: 315 Ligyrus maximus Arrow 1913: 466 Ligyrus latus Arrow 1914: 273 | Mexico, Guatemala, Honduras, Nicaragua, El Salvador, Costa Rica, Panama, Colombia, Venezuela, Ecuador, Peru, Trinidad, Brazil, Bolivia | m, f | Endrödi 1985, Ratcliffe 2003, Escalona & Joly 2006, Ratcliffe & Cave 2006, Carvajal et al. 2011, Ratcliffe et al. 2015 |
|---|
| Tomarus burmeisteri (Steinheil, 1872) Ligyrus burmeisteri Steinheil 1872: 560 Ligyrus distinctus Fairmaire 1892: 244 | Peru, Brazil, Bolivia, Paraguay, Uruguay, Argentina, Chile | m, f | Endrödi 1985 |
|---|
| Tomarus cicatricosus (Prell, 1937) Ligyrus cicatricosus Prell 1937: 90 | Mexico, Guatemala, Costa Rica, El Salvador, Honduras, Colombia | m, f | Endrödi 1985, Ratcliffe 2003, Ratcliffe & Cave 2006, López-García et al. 2015 |
|---|
| Tomarus colombianus López-García & Gasca- Álvarez, 2014 Tomarus colombianus López-García & Gasca-Álvarez, in López-García et al. 2014: 579 | Colombia | m, f | López-García et al. 2014 |
|---|