Phylogenetic analysis of Betta coccina complex (Teleostei, Osphronemidae) from Peninsular Malaysia and Sumatra Island with descriptions of two new species Author Ding, Jiali https://orcid.org/0009-0009-1322-1666 School of Life Sciences, Nanchang University, Nanchang 330031, China Author Lei, Wen https://orcid.org/0009-0005-3180-7784 School of Life Sciences, Nanchang University, Nanchang 330031, China Author Haryono, Haryono 0000-0002-5143-6790 Research Center for Biosystematics and Evolution, National Research and Innovation Agency, Bogor 16911, Indonesia Author Shi, Wentian 0000-0003-0130-7963 The Parosphromenus Project, Lage, 32792, Germany Author Zhang, Wanchang 0000-0003-1040-2516 School of Life Sciences, Nanchang University, Nanchang 330031, China text ZooKeys 2025 2025-05-15 1238 161 181 journal article 309842 10.3897/zookeys.1238.142857 b472ef28-e032-4dd4-9578-1676cd36a4c4 67271BD2-DB9C-4274-9A95-187AD2238993

Table 1. Morphometric and meristic data of Betta iaspis sp. nov. (n = 31, MZB.26963, NCUMB.65334) from Jambi, Sumatra, and Betta mulyadii sp. nov. (n = 29, MZB.26964, NCUMB.65326) from Riau, Sumatra, Indonesia.

Morphometrics	Betta iaspis sp. nov.		Betta mulyadii sp. nov.
	Holotype	Paratype	Holotype	Paratype
		minimum – maximum (mean ± standard deviation SD)		minimum – maximum (mean ± standard deviation SD)
Standard length (mm)	22.0	17.7–23.9 (21.7 ± 1.6)	18.5	22.4–28.4 (24.9 ± 1.8)
As % Standard length
Total length	132.3	112.0–131.3 (124.9 ± 4.1)	135.1	119.1–129.6 (124.3 ± 2.6)
Predorsal length	67.7	57.8–66.1 (61.6 ± 2.0)	64.9	58.1–63.5 (60.8 ± 1.4)
Postdorsal length	22.7	21.8–32.2 (25.3 ± 2.6)	24.3	21.3–28.9 (24.1 ± 2.2)
Preanal length	45.5	39.7–47.0 (42.6 ± 1.7)	42.7	38.1–43.5 (40.2 ± 1.6)
Head length	28.6	21.6–28.8 (25.1 ± 1.6)	26.9	21.2–26.2 (23.7 ± 1.2)
Body depth at dorsal-fin origin	20.0	18.2–22.3 (19.8 ± 1.1)	16.9	18.7–21.7 (23.7 ± 1.2)
Pelvic-fin length	37.7	18.3–32.8 (24.8 ± 4.3)	31.4	19.6–36.5 (28.0 ± 4.6)
Anal-fin base length	50.9	47.9–57.8 (52.7 ± 2.0)	48.1	50.7–58.9 (54.2 ± 2.0)
Dorsal-fin base length	12.8	9.87–15.1 (11.9 ± 1.3)	13.0	9.9–15.1 (12.7 ± 1.3)
Caudal peduncle depth	12.4	11.9–15.6 (14.1 ± 0.8)	12.5	12.9–15.2 (14.1 ± 0.7)
In % Head length (mean ± SD)
Orbit diameter	27.1	25.5–34.8 (29.0 ± 2.6)	30.1	24.6–37.5 (27.9 ± 3.0)
Postorbital length	50.8	49.1–62.0 (55.2 ± 3.0)	47.8	50.9–57.8 (54.9 ± 1.9)
Interorbital distance	42.8	31.6–45.3 (39.2 ± 3.6)	41.1	34.4–46.4 (40.8 ± 3.5)
Snout length	17.5	12.0–19.0 (15.8 ± 1.7)	15.6	14.1–19.3 (16.7 ± 1.4)
MERISTICS (Total counts)		minimum – maximum		minimum – maximum
Anal-fin rays	27	27–28	27	27–29
Dorsal-fin rays	10	8–10	8	8–10
Caudal-fin rays	i-I- 4 + 5 - I-i	i-I- 4 + 5 - I-i	i-I- 4 + 5 - I-i	i-I- 4 + 5 - I-i
Pelvic-fin rays	6	6	6	6
Pectoral-fin rays	12	12–14	13	12–14
Subdorsal scales	6	5–6	5	5–6
Transverse scales	8	8	8	8
Lateral scales	30	28–30	30	29–30
Predorsal scales	21	19–21	21	19–21
Postdorsal scales	10	9–11	10	9–12