Taxonomic revision of grass frogs (Ptychadenidae, Ptychadena) endemic to the Ethiopian highlands Author Goutte, Sandra https://orcid.org/0000-0002-8939-3773 New York University Abu Dhabi, Saadiyat Island, Abu Dhabi, UAE sg5533@nyu.edu Author Reyes-Velasco *, Jacobo New York University Abu Dhabi, Saadiyat Island, Abu Dhabi, UAE Author Freilich, Xenia Department of Biology, Queens College, City University of New York, Flushing, New York, USA Author Kassie, Abeje Ethiopian Biodiversity Institute, Addis Ababa, Ethiopia & Addis Ababa University, Addis Ababa, Ethiopia Author Boissinot, Stephane New York University Abu Dhabi, Saadiyat Island, Abu Dhabi, UAE stephane.boissinot@nyu.edu text ZooKeys 2021 2021-02-11 1016 77 141 http://dx.doi.org/10.3897/zookeys.1016.59699 journal article http://dx.doi.org/10.3897/zookeys.1016.59699 1313-2970-1016-77 A462DE7344284857973CBCFD17294D15 D7F1A3CBC5AC592DADE3787D4EC62BF9

Table 1. Morphometric measurements (average +/- standard deviation) of the Ptychadena neumanni complex.

Species	Sex	N	SVL	HW	HL	SL	NS	IND	EN	IOD	ETD	TD	ED	UEW	FLL	HAL	FinDW	THL	TL	FL	Toe4DW	MTL
P. amharensis	M	6	36.9 +/- 2.7	13.9 +/- 1.4	13.2 +/- 0.6	5.5 +/- 0.3	2.8 +/- 0.2	2.9 +/- 0.3	2.5 +/- 0.3	2.1 +/- 0.2	1.2 +/- 0.2	2.9 +/- 0.3	4.2 +/- 0.6	2.7 +/- 0.4	6.8 +/- 0.3	8.3 +/- 0.5	0.6 +/- 0.1	17.4 +/- 1.2	18.8 +/- 1.3	18.9 +/- 0.9	0.6 +/- 0.1	1.6 +/- 0.3
	F	7	42.5 +/- 2	13.9 +/- 0.9	14.2 +/- 0.9	6.1 +/- 0.3	3 +/- 0.5	3.3 +/- 0.2	2.6 +/- 0.3	2.4 +/- 0.3	1.5 +/- 0.3	2.9 +/- 0.3	4.3 +/- 0.4	2.9 +/- 0.2	7.6 +/- 0.6	8.5 +/- 0.5	0.6 +/- 0.1	19.2 +/- 1.3	21.2 +/- 1.3	21.1 +/- 1.3	0.5 +/- 0.2	1.7 +/- 0.2
P. beka	M	9	37.9 +/- 2.6	14.5 +/- 1	14.2 +/- 1.3	5.7 +/- 0.4	2.9 +/- 0.4	3 +/- 0.2	2.6 +/- 0.3	2.7 +/- 0.3	1.1 +/- 0.4	2.9 +/- 0.2	4 +/- 0.5	2.6 +/- 0.4	6.7 +/- 0.5	8.3 +/- 0.5	0.6 +/- 0.1	17.6 +/- 1.2	19.5 +/- 1	19.5 +/- 1	0.6 +/- 0.1	1.7 +/- 0.2
	F	8	43.2 +/- 2.9	14.7 +/- 1.2	15.6 +/- 1.3	6.7 +/- 0.5	3.4 +/- 0.4	3.7 +/- 0.3	3.2 +/- 0.3	3.2 +/- 0.5	1.5 +/- 0.2	3.2 +/- 0.3	4.5 +/- 0.7	2.9 +/- 0.3	8 +/- 0.7	9.5 +/- 0.5	0.6 +/- 0.1	20.9 +/- 1.8	24.7 +/- 1.9	23.6 +/- 1.4	0.7 +/- 0.2	1.9 +/- 0.1
P. cooperi	M	8	45 +/- 2.7	17.7 +/- 1.1	16.5 +/- 1.5	7.3 +/- 0.5	4.2 +/- 0.5	3.3 +/- 0.4	2.9 +/- 0.4	2.9 +/- 0.5	1.7 +/- 0.3	3.6 +/- 0.4	4.9 +/- 0.5	3 +/- 0.3	9.7 +/- 1.1	11.5 +/- 1.2	0.7 +/- 0.2	22.5 +/- 2.7	26.4 +/- 1.5	25.3 +/- 1.5	0.7 +/- 0.1	1.9 +/- 0.2
	F	1	61.2	19.9	20.7	7.6	3.8	4	3.1	3.4	2.2	4.6	5.8	4.8	11.4	12.6	0.9	29.4	31.9	31.1	1	2.5
P. delphina	M	6	40.1 +/- 2.9	14.7 +/- 0.4	14.3 +/- 0.8	6.2 +/- 0.3	3 +/- 0.2	3.2 +/- 0.4	2.8 +/- 0.6	2.5 +/- 0.3	1.3 +/- 0.2	3 +/- 0.4	4.2 +/- 0.6	2.9 +/- 0.2	8.1 +/- 0.5	8.9 +/- 0.9	0.6 +/- 0.1	18.8 +/- 0.8	21.4 +/- 1.5	21.6 +/- 2.6	0.5 +/- 0	1.5 +/- 0.2
	F	6	47.2 +/- 2.3	15.9 +/- 1.4	16.3 +/- 0.8	6.9 +/- 0.4	3.3 +/- 0.3	3.4 +/- 0.2	3.2 +/- 0.2	3.2 +/- 0.3	1.6 +/- 0.2	3.4 +/- 0.3	5.2 +/- 0.4	3.3 +/- 0.3	9.3 +/- 0.7	10.1 +/- 0.6	0.6 +/- 0.1	22.9 +/- 1.9	25.9 +/- 2.2	25.7 +/- 1.6	0.7 +/- 0.1	1.8 +/- 0.1
P. doro	M	13	35.8 +/- 2.2	12.6 +/- 0.8	13.1 +/- 1.1	5.6 +/- 0.4	2.7 +/- 0.3	3.1 +/- 0.2	2.5 +/- 0.2	2.8 +/- 0.4	1.1 +/- 0.2	2.5 +/- 0.3	4 +/- 0.4	2.5 +/- 0.2	7.1 +/- 0.5	8.5 +/- 0.7	0.6 +/- 0.2	17.5 +/- 1.6	20.7 +/- 1.4	21.1 +/- 2.2	0.5 +/- 0.1	1.5 +/- 0.3
	F	6	42.4 +/- 5.7	14.2 +/- 0.9	15 +/- 0.4	6.7 +/- 0.5	3.4 +/- 0.5	3.5 +/- 0.4	2.8 +/- 0.5	3.2 +/- 0.5	1.3 +/- 0.3	2.9 +/- 0.4	4.6 +/- 0.4	3 +/- 0.4	8.9 +/- 0.9	10.4 +/- 1	0.6 +/- 0.1	22.6 +/- 2.4	26.6 +/- 1.1	26.9 +/- 1.6	0.8 +/- 0.2	1.8 +/- 0.3
P. erlangeri	M	35	34.1 +/- 2.5	11.7 +/- 1	12.2 +/- 1.3	5 +/- 0.5	2.6 +/- 0.3	2.7 +/- 0.3	2.2 +/- 0.2	2.5 +/- 0.4	1 +/- 0.2	2.4 +/- 0.3	3.5 +/- 0.4	2.4 +/- 0.4	6.3 +/- 0.6	7.2 +/- 0.7	0.6 +/- 0.1	15.8 +/- 1.3	17.9 +/- 2	18.3 +/- 1.8	0.5 +/- 0.1	1.4 +/- 0.3
	F	10	34.6 +/- 1.7	11.6 +/- 1	12.2 +/- 1.3	5.2 +/- 0.5	2.7 +/- 0.3	2.9 +/- 0.2	2.5 +/- 0.2	2.5 +/- 0.4	1 +/- 0.2	2.3 +/- 0.3	3.6 +/- 0.5	2.4 +/- 0.3	6.2 +/- 0.5	7.5 +/- 0.8	0.5 +/- 0.1	16.9 +/- 2.1	18.7 +/- 2.5	19.2 +/- 1.8	0.5 +/- 0.1	1.4 +/- 0.3
P. goweri	M	15	42.4 +/- 2.7	15.7 +/- 0.9	16.2 +/- 1.4	6.7 +/- 0.5	3.1 +/- 0.3	3.5 +/- 0.4	3.1 +/- 0.4	3.2 +/- 0.4	1.5 +/- 0.2	3.1 +/- 0.4	4.3 +/- 0.7	2.9 +/- 0.2	8.5 +/- 0.9	10.2 +/- 0.8	0.7 +/- 0.2	21.9 +/- 1.5	25.4 +/- 1.5	25.7 +/- 1.2	0.7 +/- 0.1	2 +/- 0.3
	F	6	51.2 +/- 4.2	17.6 +/- 2	18.6 +/- 2.1	7.5 +/- 0.4	3.7 +/- 0.3	3.9 +/- 0.5	3.5 +/- 0.3	3.5 +/- 0.4	1.8 +/- 0.3	3.5 +/- 0.4	5 +/- 0.5	3.3 +/- 0.2	9.8 +/- 1	11.8 +/- 1	0.7 +/- 0.1	25.7 +/- 2.7	30.4 +/- 2.3	29.4 +/- 2	0.8 +/- 0.1	2.5 +/- 0.3
P. harenna	M	4	36.3 +/- 2.3	13.6 +/- 1.2	14.2 +/- 1.6	5.8 +/- 0.3	3.1 +/- 0.2	3.4 +/- 0.5	2.4 +/- 0.3	2.4 +/- 0.5	1.2 +/- 0.4	3.1 +/- 0.6	4.8 +/- 0.6	2.8 +/- 0.5	7.6 +/- 0.4	8.1 +/- 0.5	0.6 +/- 0.1	18.4 +/- 2	20.6 +/- 1.3	20.7 +/- 1.7	0.5 +/- 0.1	1.4 +/- 0.1
	F	2	43.3 +/- 0.3	15.4 +/- 1.2	16 +/- 0.8	7 +/- 0.4	3.4 +/- 0.3	3.8 +/- 0.1	3 +/- 0.1	4.2 +/- 0.2	1.4 +/- 0.1	3.1 +/- 0.4	4.7 +/- 0.1	2.7 +/- 0.6	7.8 +/- 0.5	9.8 +/- 0.6	0.8 +/- 0.4	21.8 +/- 0.4	24.9 +/- 0.4	24 +/- 0.5	0.8 +/- 0.3	2 +/- 0.5
P. levenorum	M	8	34.3 +/- 2.3	11.7 +/- 1.3	11.9 +/- 1.1	4.6 +/- 0.5	2.4 +/- 0.3	2.4 +/- 0.3	1.8 +/- 0.4	2.2 +/- 0.3	1 +/- 0.3	2.2 +/- 0.2	3.2 +/- 0.4	2.2 +/- 0.4	6.5 +/- 1	7 +/- 0.7	0.6 +/- 0.1	14.2 +/- 1.4	16.9 +/- 1.8	18 +/- 2	0.5 +/- 0.1	1.6 +/- 0.2
	F	4	37 +/- 2.5	12.2 +/- 0.9	12.1 +/- 0.5	4.8 +/- 0.5	2.5 +/- 0.3	2.7 +/- 0.4	2.1 +/- 0.2	2.6 +/- 0.3	1.3 +/- 0.2	2.4 +/- 0.4	3.5 +/- 0.6	2.5 +/- 0.4	6.6 +/- 0.8	7.5 +/- 0.5	0.5 +/- 0.2	16 +/- 1.6	18.9 +/- 2.1	20.1 +/- 2.4	0.6 +/- 0.1	1.8 +/- 0.2
P. nana	M	12	28.3 +/- 3.3	10.2 +/- 1.4	10.1 +/- 0.7	4.2 +/- 0.4	2.2 +/- 0.2	2.3 +/- 0.2	1.8 +/- 0.2	2.2 +/- 0.3	0.9 +/- 0.1	1.8 +/- 0.2	2.9 +/- 0.5	2 +/- 0.3	5 +/- 0.6	5.6 +/- 0.6	0.5 +/- 0.1	11.7 +/- 1.4	12.7 +/- 0.9	13.8 +/- 1	0.5 +/- 0.1	1.2 +/- 0.2
	F	12	29.6 +/- 3.2	9.5 +/- 1.3	10 +/- 0.8	4.2 +/- 0.4	2.3 +/- 0.3	2.5 +/- 0.3	1.9 +/- 0.2	2 +/- 0.4	1 +/- 0.2	1.8 +/- 0.2	3.2 +/- 0.4	2.1 +/- 0.4	4.9 +/- 0.3	5.8 +/- 0.6	0.5 +/- 0.1	12.2 +/- 1.2	12.9 +/- 1.1	14.3 +/- 1.2	0.5 +/- 0.1	1.2 +/- 0.2
P. neumanni	M	20	34.6 +/- 2	12 +/- 1.2	13.6 +/- 0.8	5.6 +/- 0.4	2.8 +/- 0.4	3.2 +/- 0.3	2.6 +/- 0.3	3.1 +/- 0.3	1 +/- 0.1	2.7 +/- 0.4	3.9 +/- 0.2	2.3 +/- 0.4	6.8 +/- 0.4	8.1 +/- 0.6	0.6 +/- 0.2	17.9 +/- 1.2	20.1 +/- 1.1	20.6 +/- 1.4	0.6 +/- 0.1	1.4 +/- 0.2
	F	8	45.6 +/- 1.4	15.2 +/- 1.1	16.7 +/- 1.6	7.1 +/- 0.4	3.6 +/- 0.1	4 +/- 0.4	3.2 +/- 0.3	3.6 +/- 0.4	1.5 +/- 0.3	3.4 +/- 0.4	4.7 +/- 0.5	3 +/- 0.4	8.3 +/- 0.7	9.8 +/- 0.7	0.7 +/- 0.2	21.8 +/- 1.5	26.1 +/- 0.8	26.4 +/- 1.4	0.7 +/- 0.2	1.9 +/- 0.3
P. robeensis	M	13	30.3 +/- 0.7	10.7 +/- 0.7	11.1 +/- 0.8	4.3 +/- 0.2	2.2 +/- 0.2	2.4 +/- 0.2	1.9 +/- 0.1	2.4 +/- 0.3	1.1 +/- 0.1	2 +/- 0.2	3 +/- 0.4	2.1 +/- 0.2	5.4 +/- 0.5	6.4 +/- 0.4	0.6 +/- 0.2	12.4 +/- 0.8	13.6 +/- 0.8	15.1 +/- 1	0.5 +/- 0.1	1.2 +/- 0.3
	F	4	31.6 +/- 2.9	11.1 +/- 0.6	10.9 +/- 0.2	4.3 +/- 0.4	2.2 +/- 0.5	2.5 +/- 0.2	2 +/- 0.1	2.5 +/- 0.4	1.1 +/- 0.2	2 +/- 0.2	3.3 +/- 0.1	2 +/- 0.3	5.5 +/- 0.5	6.5 +/- 0.5	0.5 +/- 0.1	12.2 +/- 1.1	13.9 +/- 0.8	15.4 +/- 1.2	0.6 +/- 0.1	1.3 +/- 0.2