Two new frog species from the Litoria rubella species group from eastern Australia Author Rowley, J. J. L. Australian Museum Research Institute, Australian Museum, 1 William St, Sydney 2010, Australia. & Centre for Ecosystem Science, School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, NSW 2052, Australia. Author Mahony, M. J. 0000-0002-1042-0848 School of Environmental and Life Sciences, The University of Newcastle, University Drive, Callaghan, 2308, Australia. michael. mahony @ newcastle. edu. au; https: // orcid. org / 0000 - 0002 - 1042 - 0848 michael.mahony@newcastle.edu.au Author Hines, H. B. Department of Environment and Science, PO Box 64, Bellbowrie, Qld, 4070, Australia. & Honorary Research Fellow, Biodiversity, Queensland Museum, PO Box 3300, South Brisbane, QLD 4101. Author Myers, S. South Australian Museum, North Terrace, Adelaide, Adelaide, 5000, Australia. & ALS Water Resources Group, 22 Dalmore Drive, Scoresby, Victoria, 3179, Australia. Author Price, L. C. School of Biological Sciences, University of Adelaide, 5005, Australia. Author Shea, G. M. Australian Museum Research Institute, Australian Museum, 1 William St, Sydney 2010, Australia. & Sydney School of Veterinary Science B 01, University of Sydney, NSW 2006, Australia. Author Donnellan, S. C. South Australian Museum, North Terrace, Adelaide, Adelaide, 5000, Australia. text Zootaxa 2021 2021-11-22 5071 1 1 41 journal article 3315 10.11646/zootaxa.5071.1.1 f94b8e6d-d8c9-4f3d-9460-9a0ffeb21a1a 1175-5326 5723255 E695DE38-387E-41E0-8188-532A907C3BB1

TABLE 1. Specimens of Litoria examined for molecular genetic analyses. Locality #—number used to identify the locality in tables and figures. SNP cluster: b— L. balatus sp. nov., d— L. dentata, and q— L. quiritatus sp. nov. ND4 —GenBank accession number. Supplementary entries in ND4 column refers to sequences <200bp in the alignment presented in the Supplementary File. ABTC Number—tissue accession number, Australian Biological Tissue Collection South Australian Museum. Voucher institutions: AMS—Australian Museum, Sydney; ANWC—Australian National Wildlife Collection, CSIRO Canberra; SAMA—South Australian Museum, Adelaide; QM—Queensland Museum, Brisbane. NP—National Park. NSW—New South Wales, Qld—Queensland.

Taxon	SNP cluster	ND4	ABTC Number	Locality #/ Locality	State	Latitude	Longitude	Voucher Institution	Voucher RegNum
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quiritatus	-	MW885785	12646	1-10 km S Mittagong, on Kangaloon Rd	NSW	-34.530	150.480	SAMA	R40858
quiritatus	-	MW885786	145056	2-Coalcliff	NSW	-34.247	150.975	AMS	R185759
quiritatus	q	KM199723	86208	3-Homebush Bay	NSW	-33.842	151.058	AMS	R147041
quiritatus	-	MW885782	86206	4-Cox’s River, Mckane’s Bridge	NSW	-33.552	150.120	AMS	R148134
quiritatus	q	MW885783	1227	5-Ourimbah	NSW	-33.340	151.330	AMS	R133185
quiritatus 5	-	KM199719	1244	5-Ourimbah	NSW	-33.340	151.330	AMS	R132687
quiritatus	-	MW885784	101713	6-Boarding House Dam	NSW	-33.001	151.404	SAMA	R69896
quiritatus	-	MW885787	101717	6-Boarding House Dam	NSW	-33.001	151.404	SAMA	R69897
quiritatus	-	MW885791	90470	6-near Sawmill Pond, Watagan National Park	NSW	-33.085	151.355	SAMA	R68522
quiritatus	-	MW885790	90497	7-Frying Pan Creek, Chichester State Forest	NSW	-32.330	151.760	SAMA	R66161
quiritatus	q	KM199720	25236	8-Mt Royal	NSW	-32.210	151.300	SAMA	R69882
dentata	d	MW885811	104273	9-Taree	NSW	-31.900	152.520	-	-
quiritatus	q	KM199719	25254	10-Mernot State Forest	NSW	-31.687	151.570	-	-
quiritatus		-	25272	10-Mernot State Forest	NSW	-31.687	151.570	-	-
quiritatus	q	MW885789	25273	10-Mernot State Forest	NSW	-31.687	151.570	-	-
dentata	-	KM199718	-	11-Lord Howe Island	NSW	-31.529	159.069	-	-
dentata	-	MW885812	123771	12-Upper Pappinbarra	NSW	-31.335	152.466	-	-
dentata	d	Supplementary	141478	12-Upper Pappinbarra	NSW	-31.335	152.467	SAMA	R69965
dentata	d	MW885802	141479	12-Upper Pappinbarra	NSW	-31.335	152.467	-	-
dentata	d	MW885809	25207	13-Hastings Forest Highway	NSW	-31.200	152.400	SAMA	R69883
dentata	d	MW885807	141458	14-Oxley Wild Rivers National Park	NSW	-30.813	152.084	AMS	R184704
dentata	d	MW885801	141459	14-East Kunderang Station, Oxley Wild Rivers National Park	NSW	-30.817	152.145	AMS	R184710
dentata	d	-	128585	15-Newry State Forest, 30 km SSW Coffs Harbour	NSW	-30.530	152.950	-	-
dentata	-	MW885816	128586	15-Newry State Forest, 30 km SSW Coffs Harbour	NSW	-30.530	152.950	-	-
dentata	-	MW885800	128587	15-Newry State Forest, 30 km SSW Coffs Harbour	NSW	-30.530	152.950	-	-
dentata	-	Supplementary	128588	15-Newry State Forest, 30 km SSW Coffs Harbour	NSW	-30.530	152.950	-	-
dentata	-	Supplementary	128582	16-Dorrigo, public camping area	NSW	-30.350	152.770	-	-
dentata	-	MW885815	128583	16-Dorrigo, public camping area	NSW	-30.350	152.770	-	-
dentata	-	MW885806	128584	16-Dorrigo, public camping area	NSW	-30.350	152.770	-	-
dentata	-	KM199716	-	17-Coffs Harbour Racecourse, Coffs Harbour	NSW	-30.308	153.127	-	-
dentata	-	KM199712	-	18-Bananacoast Caravan Park, Coffs Harbour	NSW	-30.269	153.133	-	-
dentata	-	KM199717	-	18-Bananacoast Caravan Park, Coffs Harbour	NSW	-30.269	153.133	-	-
dentata	-	KM199717	-	18-Bananacoast Caravan Park, Coffs Harbour	NSW	-30.269	153.133	-	-
dentata	d	KM199725	86359	19-between Coutts crossing and Glenreagh, N Coffs Harbour	NSW	-29.928	152.917	ANWC	A01777
dentata	-	KM199724	86356	19-Glenreagh, N Coffs Harbour	NSW	-30.050	152.983	ANWC	A01757
dentata	d	MW885803	25456	20-Mann River Nature Reserve	NSW	-29.710	152.090	SAMA	R69885
dentata	-	KM199721	25457	20-Mann River Nature Reserve	NSW	-29.710	152.090	SAMA	R69894
dentata	-	KM199705	-	21-McKittrick Park, Grafton	NSW	-29.708	152.936	-	-
dentata	-	KM199712	-	21-McKittrick Park, Grafton	NSW	-29.708	152.936	-	-
dentata	-	KM199713	-	21-McKittrick Park, Grafton	NSW	-29.708	152.936	-	-
dentata	-	KM199714	-	21-McKittrick Park, Grafton	NSW	-29.708	152.936	-	-
dentata	-	KM199715	-	21-McKittrick Park, Grafton	NSW	-29.708	152.936	-	-
dentata	-	KM199707	-	21-McKittrick Park, Grafton	NSW	-29.708	152.936	-	-
dentata	d	KM199705	86207	22-Grafton, 3.5 km N Grafton on Casino Road	NSW	-29.666	152.938	AMS	R151836
dentata	-	KM199709	-	23-Palmers Island, Yamba	NSW	-29.420	153.287	-	-
dentata	-	KM199710	-	23-Palmers Island, Yamba	NSW	-29.420	153.287	-	-
dentata	-	KM199710	-	23-Palmers Island, Yamba	NSW	-29.420	153.287	-	-
dentata	-	KM199711	-	23-Palmers Island, Yamba	NSW	-29.420	153.287	-	-
dentata	-	KM199712	-	23-Palmers Island, Yamba	NSW	-29.420	153.287	-	-
dentata	-	KM199715	-	23-Palmers Island, Yamba	NSW	-29.420	153.287	-	-
dentata	-	KM199705	-	24-Lighthouse Beach, Ballina	NSW	-28.869	153.590	-	-
dentata	-	KM199706	-	24-Lighthouse Beach, Ballina	NSW	-28.869	153.590	-	-
dentata	-	KM199705	-	25-Ballina Bowl, Ballina	NSW	-28.855	153.560	-	-
dentata	-	KM199705	-	25-Ballina Bowl, Ballina	NSW	-28.855	153.560	-	-
dentata	-	KM199707	-	25-Ballina Bowl, Ballina	NSW	-28.855	153.560	-	-
dentata	-	KM199708	-	25-Ballina Bowl, Ballina	NSW	-28.855	153.560	-	-
dentata	d	KM199722	25754	26-Peacock Creek, Richmond Range State Forest	NSW	-28.660	152.710	SAMA	R69884
dentata	d	KM199705	86376	27-Dorroughby Education Centre, Tweed Valley	NSW	-28.650	153.350	ANWC	A02008
dentata	d	MW885799	141460	28-Kyogle	NSW	-28.618	153.002	AMS	R184779
dentata	-	KM199707	25731	29-Whian Whian State Forest	NSW	-28.590	153.330	SAMA	R69895
dentata	d	KM199705	86205	30-Brunswick Heads	NSW	-28.533	153.550	AMS	R133486
dentata	d	KM199705	24826	31-Midginbil Hill	NSW	-28.502	153.262	SAMA	R40185
dentata	d	MW885805	90537	32-Border Ranges National Park	NSW	-28.400	153.030	SAMA	R68578
dentata	-	MW885813	145051	33-Headwaters of Thane Creek, Durikai State Forest	Qld	-28.288	151.696	QM	J96339
dentata	-	MW885814	145054	34-Junction of Bilborough Court and Springbrook Road, Springbrook	Qld	-28.226	153.282	QM	J96344
balatus	-	MW885796	145053	35-Junction of Upper Logan and Flanagan Reserve Roads, Barney View	Qld	-28.225	152.768	QM	J96341
dentata	-	MW885810	100638	36-between 3 and 6 km from Goomolahra Lookout, Springbrook	Qld	-28.201	153.271	QM	J86657
balatus	-	MW885795	145052	37-Boonah- Rathdowney Road, about 0.5 km E Burnett Creek Road, Maroon	Qld	-28.172	152.689	QM	J96340
dentata	-	MW885808	145055	38-Nerang- Murwillumbah Road, Numinbah Valley	Qld	-28.153	153.225	QM	J96347
balatus	b	Supplementary	127795	39-Ormeau cemetery, Ormeau	Qld	-27.805	153.267	QM	J92764
balatus	b	Supplementary	127796	39-Ormeau cemetery, Ormeau	Qld	-27.805	153.267	QM	J92765
balatus	-	Supplementary	127799	40-Junction of Wharf Road and Pimpama, Jacobs Creek Road, Pimpama	Qld	-27.800	153.293	QM	J92768
balatus	-	Supplementary	127801	41-Woongoolba Conservation Park, Woongoolba	Qld	-27.746	153.308	QM	J92770
balatus	-	Supplementary	127802	41-Woongoolba Conservation Park, Woongoolba	Qld	-27.746	153.308	QM	J92771
balatus	-	-	100641	42-Karawatha Forest Park	Qld	-27.634	153.095	QM	J86660
balatus	b	MW885793	100642	42-Karawatha Forest Park	Qld	-27.634	153.095	QM	J86661
balatus	-	-	100643	42-Karawatha Forest Park	Qld	-27.634	153.095	QM	J86662
balatus	-	Supplementary	127707	43-junction of Moggill Road and Aitcheson Street, Moggill	Qld	-27.587	152.860	QM	J92496
balatus	-	MW885797	127708	43-junction of Moggill Road and Aitcheson Street, Moggill	Qld	-27.587	152.860	QM	J92497
balatus	b	-	110165	44-Anstead Bushland Reserve	Qld	-27.587	152.860	QM	J91105
balatus	-	Supplementary	110166	44-Anstead Bushland Reserve	Qld	-27.541	152.861	QM	J91106
balatus	b	MW885792	127658	45-Esk- Hampton Road, Perseverance	Qld	-27.373	152.108	QM	J93230
balatus	b	Supplementary	127659	45-Esk- Hampton Road, Perseverance	Qld	-27.373	152.108	QM	J93231
balatus	-	MW885794	127663	46-156 Zillman Road, Ocean View	Qld	-27.125	152.818	QM	J93232
balatus	b	Supplementary	141466	47-Mount Victoria property, adjoining Bunya Mountains NP	Qld	-26.929	151.633	QM	J95155
balatus	b	Supplementary	141467	47-Mount	Qld	-26.929	151.633	QM	J95156
				Victoria property, adjoining Bunya Mountains NP
balatus	-	MW885798	145050	48-West Kilcoy Creek, near to Conondale NP	Qld	-26.785	152.563	QM	J96338
congenita	79204	MW885823	79204	Aquam Camp, Trans-Fly, Papua New Guinea	Western	-	-	-	-
congenita	-	MW885817	-	Nenas, West Timor, Indonesia	East Nusa Tenggara	-9.55	124.21	WAM	R106728
congenita	79184	-	79184	Wegamu, Trans- Fly, Papua New Guinea	Western	-8.4328	141.1128	SAMA	R69842
electrica	-	MW885780	13053	Cumberland Dam, 20 km W Georgetown	Qld	-18.30	143.350	-	-
electrica	-	MW885781	16519	Burketown	Qld	-17.75	139.55	SAMA	R33970
electrica	-	KM199726	16521	Burketown	Qld	-17.75	139.55	SAMA	R33972
pygmaea	-	100661	100661	Wondiwoi Mountains, Indonesia	West Papua	-	-	ZMB	70519
pygmaea	-	100665	100665	6 km N Kontiunae, Waira Mt, Yapen Island, Indonesia	West Papua	-	-	ZMB	62323
rubella	-	MW885777	100006	Mt Brockman	WA	-	-	WAM	R135374
rubella	127683	MW885778	127683	Larcom Creek, S Mount Larcom	Qld	-23.871	151.016	QM	J90515
rubella	-	MW885779	135591	Camballin	WA	-17.982	124.2	WAM	R140793
rubella	-	KM199727	1233	Collaroy Homestead	NSW	-31.466	146.65	AMS	R132676
capitula	-	MW885779	135591	Latdalam, Tanimbar, Indonesia	Maluku	-7.983	131.150	WAM	R109984
capitula	135592	-	135592	Latdalam, Tanimbar, Indonesia	Maluku	-7.983	131.150	WAM	R109985
amboinensis	-	MW885825	100658	Wondiwoi Mountains, Indonesia	West Papua	-	-	ZMB	62090
darlingtoni	-	MW885822	43111	Yuro, Papua New Guinea	Chimbu	-6.533	144.85	AMS	R114601
everetti	-	MW885817	-	Nenas, West Timor, Indonesia	East Nusa Tenggara	-9.55	124.21	WAM	R106728
everetti	-	MW885818	-	Baa, Roti Island, Indonesia	East Nusa Tenggara	-10.73	123.1	WAM	R105676
peronii	-	MW885820	3885	Cooyar Creek crossing, E Yarraman	Qld	-26.87	152.03	SAMA	R33658
peronii	-	MW885818	24187	Ourimbah	NSW	-33.37	151.37	SAMA	R33915
rothii	-	MW885819	1082	Pinnacles	Qld	-15.68	143.56	SAMA	R32540
rothii	-	MW885824	100095	Mount Elizabeth Station	WA	-16.419	126.100	WAM	R167959