From folkloric origins to scientific systematics: the first detailed redescription of the type species of Lycosa Latreille, 1804 with insight into the genus (Araneae: Lycosidae) Author Ballarin, Francesco Systematic Zoology Laboratory, Department of Biological Sciences, Tokyo Metropolitan University, 1 – 1 Minami-Osawa, Hachioji-shi, 192 - 0397, Tokyo, Japan & Department of Zoology, Museo di Storia Naturale of Verona, Lungadige Porta Vittoria, 9, I- 37129, Verona, Italy ballarin.francesco@gmail.com Author Marusik, Yuri M. Institute for Biological Problems of the North, FEB RAS, Portovaya Str., 18, RF- 685000, Magadan, Russia & Department of Zoology and Entomology, University of the Free State, 9300, Bloemfontein, South Africa yurmar@mail.ru Author Pantini, Paolo Museo Civico di Scienze Naturali “ Enrico Caffi ”, Piazza Cittadella, 10, I- 24129, Bergamo, Italy paolo.pantini@comune.bergamo.it Author Nadolny, Anton A. A. O. Kovalevsky Institute of Biology of the Southern Seas, Nakhimov Ave., 2, Sevastopol, Crimea nadolnyanton@ibss-ras.ru text European Journal of Taxonomy 2026 2026-03-17 1043 244 286 https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3221/14275 journal article 314828 10.5852/ejt.2026.1043.3221 040d868b-9b35-4c6c-9ad7-d885cc309426 2118-9773 19224663 31AB292F-9FFF-4DBF-8FBC-BF05C05BE99E

Table 5 (continued). Provisional members of Lycosa s. str. based on currently available morphological and molecular information.

			Additional references
	Known		other than the original
Species names	sex	Distribution	description
L. piochardi	♂♀	Turkey, Egypt, Israel,	Armiach Steinpress et
Simon, 1876		Palestine, Jordan,	al. 2022; Al-Khazali et
		Lebanon, Syria, Iraq,	al. 2023; Logunov 2023.
		Iran	See also WSC 2025 and
			Nentwig et al. 2025
L. praegrandis	♂♀	Albania, North	Zyuzin & Logunov
C. L. Koch, 1836		Macedonia, Bulgaria,	2000; Armiach
		Greece, Turkey, Levant,	Steinpress et al. 2022;
		Ukraine (south part near	Logunov 2023; Esyunin
		the seas), Russia (south	et al. 2024. See also
		of European part),	WSC 2025 and Nentwig
		Azerbaijan, Armenia,	et al. 2025
		Kazakhstan, Uzbekistan,
		Turkmenistan,
		Tajikistan, Iran
L. soboutii	♂♀	Georgia, Iran	Logunov 2023; Seropian
Shafaie, Nadolny & Mirshamsi, 2022			et al. 2024
L. suboculata	♀	Morocco	Planas et al. 2013
Guy, 1966
L. tarantula	♂♀	South France, Italy	Pepe 2005; Planas et
(Linnaeus, 1758)		mainland (central-	al. 2013; this study. See
		south), Slovenia,	also WSC 2025 and
		Croatia. Questionable	Nentwig et al. 2025
		records are discussed in
		the text
L. uzbekistanica	♀	Uzbekistan	–
Logunov, 2023
L. vachoni	♀	Algeria, Tunisia	Planas et al. 2013
Guy, 1966
L. wadaiensis	♀	Chad	–
Roewer, 1960