Morphological and phylogenetic analysis approach to three new species and a new section of Astragalus (Fabaceae) from Mongolia Author Munkhtulga, Dariganga https://orcid.org/0009-0009-0175-203X Department of Biology, School of Arts and Sciences, National University of Mongolia, Ulaanbaatar 14201, Mongolia Author Baasanmunkh, Shukherdorj 0000-0003-4224-9376 Department of Biology and Chemistry, Changwon National University, Changwon 51140, Korea Author Nyamgerel, Nudkhuu 0000-0002-9110-423X Department of Biology and Chemistry, Changwon National University, Changwon 51140, Korea Author Park, Jong Ho https://orcid.org/0009-0003-9053-8591 Department of Biology and Chemistry, Changwon National University, Changwon 51140, Korea Author Tsegmed, Zagarjav 0000-0002-6024-4918 Department of Biology and Chemistry, Changwon National University, Changwon 51140, Korea Author Tojibaev, Komiljon Sh. 0000-0003-2846-5777 Institute of Botany, Academy of Sciences of Uzbekistan, Tashkent 100125, Uzbekistan Author Choi, Hyeok Jae 0000-0001-6315-0071 Department of Biology and Chemistry, Changwon National University, Changwon 51140, Korea text PhytoKeys 2025 2025-04-08 255 51 73 journal article 309089 10.3897/phytokeys.255.140805 1884c5d7-82ed-4f76-90c5-97ebfd919f60

Table 2. Morphological comparisons of five Astragalus species in Mongolia.

Characters	A. laguroides	A. gobi-altaicus	A. oyunicus	A. teshigicus	A. uvsicus
Leaves	3-14 (- 18) cm long	8-14 cm long	5-12 (- 15) cm long	8-18 cm long	3-8 (- 12) cm long
Leaflets	3-5 (- 8) pairs, narrowly elliptic, 8-25 (- 40) × 3-6 mm, both surfaces rather densely hairy, apex acute	2-4 pairs, broadly-elliptical or oblong-obovate, 7-12 × 3–5 mm, on both sides densely hairy, apex mucronate	3-5 pairs, narrowly elliptic, 15-18 × 4-7 mm, on both sides densely hairy, apex acute	4-6 pairs, elliptical, 14-20 × 5-8 mm, on both sides densely covered with medifixed, subappressed hairs, acute to rarely obtuse	2-5 pairs, narrowly elliptic, 8-11 (- 14) × 3-5 mm, rather densely covered with medifixed, appressed hairs, apex acute
Stipules	5-8 (- 10) mm long, narrowly triangular-acuminate, covered with spreading or ascending hairs	3-4 mm long, narrowly triangular-acuminate, rather densely appressed hairy, at the margins with basifixed hairs	4-6 mm long, broadly triangular-acuminate, loosely covered with spreading or ascending hairs	4-5 mm long, broadly triangular-acuminate, densely covered with spreading or ascending hairs	6-8 mm long, narrowly triangular-acuminate, covered with spreading or ascending hairs
Raceme	ovoid to oblong, 2-4 (- 6) cm long	oblong-cylindrical, 3-8 cm long	ovoid to oblong, 3-4 cm long	oblong or oblong-cylindrical, 4-7 (- 8) cm long	globose or ovate, 3-4 cm long
Calyx	11-14 mm long, densely covered with asymmetrically bifurcate to basifixed, tangled, spreading white hairs 1–2 mm, at outer side of the teeth and sometimes at the nerves of the tube also with black hairs; teeth 3-4 mm	12-16 mm long, densely covered with subbasifixed to basifixed, spreading, straight white hairs 2-3 mm; teeth 4-6 mm	10-12 mm long, covered with basifixed, spreading, straight white and fewer black hairs 1-2 mm; teeth 2-3 mm	12-15 mm long, rather densely covered with spreading, straight white hairs 2 mm; teeth 2-3 mm	11-14 mm long, covered with basifixed, spreading white and black hairs 1-2 mm; teeth 2-3 mm
Standard	17-20 mm long, emarginate	14-18 (- 20) mm long, slightly emarginate	18-20 mm long, emarginate	20-24 mm long, emarginate	15-18 mm long, emarginate
Wings	14-18 mm long	12-16 mm long	14-18 mm long	17-22 mm long	13-15 mm long, obtuse
Keels	13-15 mm long	10-14 mm long	12-16 mm long	14-18 mm long	12-14 mm long
Pods	sessile, oblong, (- 6) 7-8 × 2-3 mm, with a hooked beak c. 1 mm, unilocular; covered with subappressed to spreading white hairs 1-1.5 mm, smaller black hairs amount greater than the apex	sessile, oblong, 7-8 × 3-4 mm, with a hooked beak 2-2.5 mm, unilocular; covered with tangled, spreading white hairs	sessile, oblong, 6-7 × 3-4 mm, with a hooked beak c. 1 mm, unilocular; densely covered with spreading white and fewer black hairs c. 2 mm	sessile, oblong, 5-6 × 2-3 mm, with a hooked beak c. 1 mm, unilocular; densely covered with spreading white hairs	sessile, linear, 6.5-7 × 2-3 mm, with beak c. 1 mm, bilocular, loosely covered with spreading white hairs c. 1 mm, smaller black hairs amount greater than the apex