A taxonomic revision of the whitefish of lakes Brienz and Thun, Switzerland, with descriptions of four new species (Teleostei, Coregonidae) Author Selz, Oliver M. Department of Fish Ecology and Evolution, Centre for Ecology, Evolution & Biogeochemistry, Eawag: Swiss Federal Institute of Aquatic Science and Technology, 6047 Kastanienbaum, Switzerland & Aquatic Ecology and Evolution, Institute of Ecology and Evolution, University of Bern, 3012 Bern, Switzerland https://orcid.org/0000-0002-2210-5909 oliver.selz@eawag.ch Author Doenz, Carmela J. Department of Fish Ecology and Evolution, Centre for Ecology, Evolution & Biogeochemistry, Eawag: Swiss Federal Institute of Aquatic Science and Technology, 6047 Kastanienbaum, Switzerland & Aquatic Ecology and Evolution, Institute of Ecology and Evolution, University of Bern, 3012 Bern, Switzerland Author Vonlanthen, Pascal Department of Fish Ecology and Evolution, Centre for Ecology, Evolution & Biogeochemistry, Eawag: Swiss Federal Institute of Aquatic Science and Technology, 6047 Kastanienbaum, Switzerland & Aquabios GmbH, Les Fermes 57, 1792 Cordast, Switzerland Author Seehausen, Ole Department of Fish Ecology and Evolution, Centre for Ecology, Evolution & Biogeochemistry, Eawag: Swiss Federal Institute of Aquatic Science and Technology, 6047 Kastanienbaum, Switzerland & Aquatic Ecology and Evolution, Institute of Ecology and Evolution, University of Bern, 3012 Bern, Switzerland text ZooKeys 2020 989 79 162 http://dx.doi.org/10.3897/zookeys.989.32822 journal article http://dx.doi.org/10.3897/zookeys.989.32822 1313-2970-989-79 F78F6D879DDB4CD98E4C60E4883A59B6 65AB71A5B3985F5E9FBF8CD56BC96143

Table 12. Morphological and meristic data of C. gutturosus Gmelin, 1818, C. arenicolus Kottelat, 1997, C. macrophthalmus Nuesslin, 1882 and C. wartmanni Bloch, 1784 from Lake Constance. Coregonus gutturosus Gmelin, 1818, non-types N = 10. Coregonus arenicolus Kottelat, 1997, holotype, NMBE-1076223 (Eawag-239-1), sex unknown; paratypes N = 3. Coregonus macrophthalmus Nuesslin, 1882, syntypes N = 7. C. wartmanni Bloch, 1784, non- type, NMBE-1076206, female.

Morphological characters	C. gutturosus			C. arenicolus				C. macrophthalmus			C. wartmanni
	Non-types (N=10)			Holotype	Paratypes (N=3)			Syntypes (N=7)			Non-type
	N -total	Mean +/- StDev	Range		N -total	Mean +/- StDev	Range	N -total	Mean +/- StDev	Range
SL (mm)	10	220.4 +/- 36.8	(169-292)	296.0	3	301.3 +/- 12.5	(289-314)	7	213.9 +/- 12.4	(193-235)	301
Percentage of standard length
PelvFB	10	4.1 +/- 0.2	(3.7-4.4)	3.9	3	4.4 +/- 0.3	(3.9-4.6)	7	3.8 +/- 0.3	(3.3-4.2)	3.8
PelvFS	10	6.1 +/- 0.4	(5.3-6.7)	5.4	3	5.7 +/- 0.5	(5.2-6.1)	7	5.7 +/- 0.7	(4.8-6.9)	6.5
PelvF	10	17.1 +/- 1.2	(15.4-19.1)	17.3	3	17.3 +/- 0.7	(16.8-18.1)	7	16.5 +/- 0.9	(15.2-17.6)	15.4
PecFB	10	3.4 +/- 0.3	(2.9-3.9)	3.3	3	3.4 +/- 0.2	(3.2-3.5)	7	3.2 +/- 0.4	(2.8-3.9)	3
PecF1	10	16.8 +/- 1.1	(14.8-18.9)	14.4	3	16.8 +/- 0.5	(14.4-17.2)	7	16.4 +/- 1.2	(15.1-18.1)	16
PecF2	10	18.2 +/- 1.5	(16.8-20.3)	15.7	3	17.4 +/- 0.8	(15.7-18)	7	17.1 +/- 1	(15.6-18.4)	17
DFB	10	11.9 +/- 0.7	(10.7-12.8)	12.2	3	12.2 +/- 1.1	(11.0-13.1)	7	11.6 +/- 0.7	(10.8-12.4)	11.2
DFAe	10	19.3 +/- 1.3	(17.6-21.6)	18.9	3	19.2 +/- 1.2	(18.0-20.3)	7	18.2 +/- 1.2	(16.6-19.6)	16.6
DFAd	10	20.4 +/- 1.1	(19.0-22.2)	20.2	3	20.5 +/- 1.3	(19.3-21.9)	7	19.2 +/- 1.3	(17.2-20.5)	18.2
DFPe	10	5.5 +/- 0.7	(4.8-7.0)	5.2	3	5.5 +/- 0.2	(5.2-5.7)	7	5.2 +/- 0.6	(4.4-5.9)	4.6
AFB	10	12.4 +/- 0.8	(11.4-13.4)	11.9	3	11.5 +/- 1.1	(10.7-12.7)	7	12.3 +/- 1.3	(10.6-14.2)	12.5
AFAe	10	12.3 +/- 1.0	(10.7-13.9)	13.2	3	13.3 +/- 0.5	(12.9-13.8)	5	12.1 +/- 1.3	(10.8-13.9)	11.1
AdFB	10	5.6 +/- 0.4	(4.9-6.1)	5.7	3	5.0 +/- 1.3	(3.7-6.2)	7	5.3 +/- 0.3	(4.9-5.8)	4
CF	9	23.2 +/- 1.9	(20.8-25.6)	na	2	24 +/- 0.1	(24-24.1)	3	22.6 +/- 1.2	(21.8-24)	23.8
CD	10	7.4 +/- 0.4	(6.7-8.2)	7.7	3	8.1 +/- 0.1	(7.7-8.2)	7	7.4 +/- 0.4	(6.9-8)	7.4
CL	10	12.9 +/- 0.8	(11.5-13.9)	14.4	3	12.9 +/- 0.8	(12.0-14.4)	7	13.8 +/- 1.3	(12.4-16.5)	13.2
PAdC	10	18.5 +/- 0.6	(17.4-19.3)	19.6	3	17.2 +/- 2.2	(14.6-19.6)	7	18.9 +/- 1.1	(17.6-20.2)	17.8
DHL	10	16.8 +/- 0.8	(15.4-18.1)	15.1	3	15.1 +/- 0.2	(14.8-15.3)	7	15.7 +/- 0.8	(14.4-16.5)	14.5
PreP	10	52.7 +/- 1.4	(50.4-54.1)	49.5	3	50.6 +/- 0.5	(49.5-51.0)	7	51.7 +/- 1.7	(48.1-53.1)	50.7
PreA	10	77.9 +/- 1.4	(76.0-80.4)	75.0	3	79.2 +/- 0.9	(75.0-80.3)	7	76.8 +/- 0.9	(75.7-78.3)	77.4
PreD	10	48.4 +/- 0.9	(46.8-49.6)	47.9	3	49.2 +/- 0.5	(47.9-49.6)	7	47 +/- 1	(45.8-48.5)	47.3
BD	10	25.9 +/- 1.9	(22.9-29.6)	24.4	3	26.2 +/- 0.8	(24.4-27.1)	7	23.5 +/- 1.9	(21.0-26.9)	23.5
PostD	10	43 +/- 1.5	(40.6-45.4)	44.8	3	42.2 +/- 1.7	(40.4-44.8)	7	43 +/- 1.4	(41.6-45.7)	44
TL	9	120.2 +/- 3	(115.1-124.3)	na	2	121.5 +/- 2.9	(119.4-123.5)	3	119.2 +/- 0.7	(118.9-120)	120.6
HL (mm)	10	50.3 +/- 7.1	(41.6-62.4)	61.8	3	63.7 +/- 3.9	(59.6-67.2)	7	47.5 +/- 3.2	(42.6-51.3)	64.7
Percentage of head length
SN	10	22.4 +/- 0.7	(21.1-23.1)	23.4	3	23.4 +/- 1.6	(21.6-24.6)	7	21.7 +/- 2.7	(18-25.6)	24
ED	10	21.1 +/- 1.4	(19.4-23)	19.6	3	17.7 +/- 0.4	(17.3-19.6)	7	24.1 +/- 1.7	(21.3-26.1)	18.9
EC	10	26.9 +/- 1.2	(25.4-29.3)	25.7	3	25 +/- 0.8	(24.1-25.7)	7	28.9 +/- 2	(25.4-30.8)	23.9
EH	10	21.3 +/- 0.6	(20.5-22.6)	20.8	3	19.6 +/- 0.9	(18.8-20.8)	7	23.2 +/- 2.1	(19.5-25.6)	19
ES	10	4.8 +/- 0.7	(3.5-5.6)	4.9	3	5.2 +/- 0.5	(4.6-5.5)	7	3.9 +/- 0.8	(2.7-4.6)	5.1
PostO	10	52.8 +/- 1	(51.5-54.4)	55.7	3	54 +/- 1	(53-55.7)	7	50.2 +/- 1.9	(48.5-53.2)	53.4
HD	10	74.2 +/- 3.2	(69.9-80.6)	68.2	3	72.6 +/- 3	(68.2-75)	7	68.6 +/- 4.8	(61.6-76.3)	67.6
MW	10	9.8 +/- 0.6	(9-11.2)	10.4	3	10.5 +/- 0.5	(10-11)	6	10.1 +/- 0.8	(8.7-11.1)	10.6
UJ	10	26.8 +/- 1.2	(24.6-29)	27.2	3	29.3 +/- 1	(27.2-30.1)	7	30.3 +/- 2.3	(26.7-33.8)	28.8
LJ	10	36.6 +/- 1.4	(34.3-39.1)	37.8	3	38.7 +/- 0.6	(37.8-39.1)	7	42.2 +/- 2	(40-44.4)	43.5
M	10	18.9 +/- 1.3	(17.3-21.7)	21.1	3	19.7 +/- 0.9	(18.6-21.1)	5	23.1 +/- 1.9	(20.1-24.7)	22
SD	10	10.2 +/- 0.8	(9.3-11.9)	9.7	3	10.9 +/- 1.3	(9.7-12.3)	7	7.4 +/- 1.2	(5.5-9.5)	6.8
SW	10	15.1 +/- 1.6	(12.3-17.6)	14.9	3	17.8 +/- 0.7	(14.9-18.5)	7	15.6 +/- 1.2	(14.1-17.4)	15
HW	10	56.1 +/- 4.3	(46.7-62.3)	51.8	3	50.8 +/- 0.5	(50.5-51.8)	7	41.6 +/- 1.5	(39.3-43.3)	45.5
IOW	10	28.4 +/- 1.7	(26.2-31.6)	29.6	3	29.7 +/- 1	(28.8-30.8)	7	26.1 +/- 1.6	(23.8-28.9)	24.2
INW	10	11.9 +/- 0.7	(10.7-12.7)	12.0	3	13.7 +/- 0.1	(12-13.8)	5	11.9 +/- 1.3	(10.7-14.1)	12.7
LJW	10	7.7 +/- 1	(6.8-9.9)	7.8	3	8.1 +/- 0.3	(7.8-8.5)	5	7.8 +/- 1	(6.4-8.8)	8.1
UJW	10	25.2 +/- 1.2	(23.1-26.8)	24.9	3	26.4 +/- 0.8	(24.9-27.2)	7	21.6 +/- 1.9	(18.6-24.6)	22.7
MGR	9	6.9 +/- 1.3	(4.1-8.7)	9.9	2	10.2 +/- 0.6	(9.8-10.6)	4	12.5 +/- 1.4	(11.6-14.7)	10.8
LGR	9	8.2 +/- 1.4	(6.7-10.6)	10.9	2	11.5 +/- 0.6	(10.9-12)	4	14.6 +/- 1.2	(13.3-16.1)	11.3
UA	na	na	na	na	na	na	na	na	na	na	na
LA	na	na	na	na	na	na	na	na	na	na	na
Meristic character	N -total	Mode	Range		N -total	Mode	Range	N -total	Mode	Range
PelvF nbranched	10	1	(1-1)	1	3	1	(1-1)	7	1	(1-1)	1
PelvF branched	10	11	(9-11)	11	3	11	(11-11)	7	10	(10-11)	12
PecF nbranched	10	1	(1-1)	1	3	1	(1-1)	7	1	(1-1)	1
PecF branched	10	13	(12-14)	12	3	12	(12-14)	7	15	(14-15)	16
DF unbranched	10	4	(3-4)	4	3	4	(4-4)	7	4	(4-4)	4
DF branched	10	10	(9-10)	9	3	10	(9-10)	7	9	(9-10)	10
AF unbranched	10	3	(3-4)	3	3	3	(3-3)	7	3	(3-4)	4
AF branched	10	11	(10-12)	10	3	10	(10-11)	7	11	(10-13)	13
LS	10	78	(76-82)	82	3	na	(82-90)	7	80	(73-80)	84
PDS	10	33	(31-35)	36	3	na	(36-44)	7	32	(32-36)	34
TDS	10	10	(9-10)	10	3	10	(10-11)	7	9	(9-10)	10
TAS	10	8	(7-9)	8	3	9	(8-9)	7	7	(7-9)	8
TPS	10	8	(8-9)	8	3	9	(8-9)	7	7	(7-9)	8
UGR	9	7	(7-9)	9	2	na	(9-12)	4	14	(12-14)	11
LGR	9	10	(9-12)	13	2	na	(13-19)	4	24	(22-24)	23
total GR	9	19	(16-21)	22	2	na	(22-31)	4	36	(36-38)	34