Increasing the species diversity of the monotypic genus Pariolius Cope 1872 (Siluriformes: Heptapteridae) after more than 150 years Author Faustino-Fuster, Dario R. 0000-0002-1445-3495 Departamento de Ictiología, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Lima, Perú & Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Programa de Pós-Graduação em Biologia Animal, Porto Alegre, Brasil & Grupo de Investigación Cuencas, Fundación Neotropical Cuencas, Arauca, Colombia dariorff36@gmail.com Author López-Castaño, Jeisson A. 0000-0002-1041-1271 Facultad de Ciencias Biológicas, Universidad Nacional Mayor de San Marcos, Ciudad Universitaria, Lima, Perú Author Quiñones, Jhonatan M. 0000-0001-8713-0024 Facultad de Ciencias Biológicas, Universidad Nacional Mayor de San Marcos, Ciudad Universitaria, Lima, Perú & Grupo de Investigación en Recursos Hidrobiológicos y Pesqueros GIREPHES, Universidad de los Llanos, Meta, Colombia Author Meza-Vargas, Vanessa 0009-0009-1623-2336 Departamento de Ictiología, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Lima, Perú & Grupo de Investigación Cuencas, Fundación Neotropical Cuencas, Arauca, Colombia text Zootaxa 2024 2024-04-08 5433 3 389 403 http://dx.doi.org/10.11646/zootaxa.5433.3.6 journal article 293428 10.11646/zootaxa.5433.3.6 f426a1c0-1b63-4845-a3be-e232e5b43df5 1175-5326 10954273 EDE5E5E3-9A6C-4347-BAA4-E7A0F01F52FC

TABLE 1. Morphometric data of Pariolius species. H (Holotype); Min (minimum); Max (maximum); SD (standard deviation); N= individuals.

	Pariolius armillatus Cope 1872					Pariolius pax New species						Pariolius maldonadoi New species
...Continued on the next page
	N	Min	Max	Mean	SD	N	H	Min	Max	Mean	SD	N	H	Min	Max	Mean	SD
Standard length (mm)	12	21.4	37.6	28.5	-	19	38.8	25.5	39.7	31.7	-	16	29.9	17.8	38.8	27.8	-
Percents of Standard Length
Predorsal distance	11	44.7	49.8	47.5	1.7	19	45.4	44.1	49.6	46.3	1.4	16	43.9	42.2	46.0	44.2	1.0
Preadipose distance	11	74.0	79.6	76.7	1.5	19	71.8	70.2	76.6	71.8	1.5	16	72.2	70.8	74.4	72.7	0.8
Prepectoral distance	11	21.2	26.4	23.6	1.6	19	21.4	19.2	24.8	23.1	1.3	16	23.6	19.6	26.2	23.0	1.6
Prepelvic distance	11	39.8	46.0	43.3	2.1	19	41.0	40.7	44.8	42.8	1.2	16	42.3	36.7	42.4	40.5	1.5
Preanal distance	11	66.6	74.3	70.1	2.2	19	69.1	64.1	69.1	67.5	1.1	16	69.1	61.8	70.1	67.2	1.8
Body depth	11	8.8	14.7	12.4	1.7	19	13.5	13.3	17.0	14.4	0.9	16	11.2	9.5	13.2	11.6	0.9
Caudal peduncle depth	11	7.3	9.2	8.5	0.6	19	9.8	8.5	10.4	9.5	0.5	16	8.9	7.6	9.0	8.4	0.4
Caudal peduncle length	11	11.7	15.3	13.3	1.1	19	15.6	13.2	16.5	15.0	0.9	16	14.7	14.3	17.2	15.6	0.8
Body width	6	14.3	18.5	17.0	1.6	19	17.1	15.9	18.9	17.6	0.8	16	16.2	13.7	17.9	16.3	1.0
Dorsal-fin base length	11	9.5	12.7	11.6	1.0	19	10.6	10.3	12.8	11.4	0.6	16	11.4	11.0	12.4	11.6	0.4
Anal-fin base length	11	16.4	20.1	17.6	1.2	19	18.1	16.9	18.7	18.0	0.4	16	18.1	14.7	20.8	18.1	1.3
Unbranched dorsal-fin ray length	11	10.8	19.6	14.1	2.7	19	9.9	9.3	13.6	12.4	1.3	16	12.9	9.2	14.1	12.8	1.2
Dorsal-fin length	11	11.0	19.4	15.0	2.5	19	15.6	14.3	16.8	15.4	0.8	16	16.3	11.8	18.0	15.3	1.3
Adipose-fin depth	11	2.0	3.6	2.6	0.5	19	3.1	3.1	4.6	3.8	0.4	16	2.9	1.9	4.7	3.2	0.8
Adipose-fin base length	11	17.1	21.0	19.2	1.5	19	23.6	20.8	25.1	23.6	1.0	15	23.3	22.7	25.1	22.5	0.7
Interdorsal distance	11	15.6	19.4	17.4	1.3	19	17.1	12.4	18.0	15.8	1.4	16	17.8	16.7	18.5	17.4	0.5
Unbranched pectoral-fin ray length	11	9.9	17.5	12.6	2.6	19	11.9	10.3	14.0	12.2	1.0	16	13.3	9.8	13.8	12.9	0.9
Pectoral-fin length	11	11.5	15.6	13.7	1.5	19	15.6	13.0	16.0	14.8	0.8	16	14.9	11.4	16.5	15.2	1.2
Pelvic-fin length	11	10.8	16.9	13.5	2.2	19	13.4	12.5	16.2	14.2	1.1	16	14.5	13.9	17.0	15.1	1.0
Pectoral-pelvic fins distance	11	18.4	23.6	21.3	1.6	19	20.6	20.6	24.6	22.1	1.1	16	20.0	17.3	23.0	20.0	1.2
Pelvic-anal fins distance	11	25.3	28.8	26.5	1.0	19	25.5	24.3	26.3	25.3	0.5	16	27.6	26.0	28.1	27.1	0.7
Dorsal-fin insertion-hypural plate	6	52.7	55.8	53.7	1.1	19	55.8	53.5	58.3	55.6	1.0	16	56.8	55.1	58.9	56.7	1.0
	N	Min	Max	Mean	SD	N	H	Min	Max	Mean	SD	N	H	Min	Max	Mean	SD
Pelvic-fin insertion-hypural plate	6	56.7	59.7	57.8	1.2	19	60.2	57.4	60.2	58.8	0.8	16	60.9	58.1	62.6	60.8	1.1
Anal-fin insertion-hypural plate	11	30.5	34.9	31.9	1.6	19	33.2	32.7	35.0	33.7	0.7	16	33.4	31.9	36.1	33.5	1.2
Ventral caudal-fin lobe length	11	16.3	24.0	20.4	2.2	19	18.3	17.4	21.2	19.8	0.9	15	21.5	18.0	25.8	22.5	1.8
Dorsal caudal-fin lobe length	11	18.5	25.5	22.9	2.0	19	21.9	20.7	25.7	23.3	1.2	14	29.7	24.4	42.6	29.6	4.2
Percents of Head Length
Head length	11	20.7	26.8	23.3	1.5	19	22.5	22.5	26.1	25.0	0.9	16	23.4	21.7	26.7	24.0	1.2
Snout length	11	30.2	40.2	34.8	2.5	19	33.2	32.9	37.2	35.2	1.2	16	35.9	31.4	37.8	35.3	2.1
Orbital diameter	11	6.8	15.0	11.1	2.8	19	10.0	8.5	11.5	10.0	0.7	16	11.6	3.7	12.0	10.6	1.9
Head width	11	68.3	94.2	82.3	10.0	19	72.0	69.0	73.0	71.7	0.9	16	66.3	64.3	68.3	66.3	1.2
Mouth width	11	41.5	58.8	49.5	6.5	19	46.0	42.8	50.9	46.6	1.6	16	46.2	44.3	47.7	46.0	1.1
Maxillary barbel length	11	99.3	137.3	122.2	11.6	19	93.7	83.5	104.9	95.3	7.2	16	109.8	91.9	119.4	107.8	6.5
External mandibular barbel length	11	62.8	102.4	86.3	11.7	19	74.9	66.4	79.6	74.6	3.3	16	78.5	77.5	88.2	82.7	3.4
Internal mandibular barbel length	11	51.4	71.3	57.5	5.3	19	59.5	55.7	67.5	59.9	2.7	16	60.2	55.4	66.3	59.3	3.4
Postorbital distance	6	47.7	63.4	55.4	6.6	19	57.8	56.0	60.4	58.3	1.2	16	56.1	55.3	59.7	56.9	1.2
Interorbital width	11	22.0	29.4	26.8	2.4	19	25.9	23.4	26.3	24.9	0.8	16	23.1	20.1	27.7	24.0	1.8
Snout-anterior nostril distance	6	11.5	13.7	12.8	0.8	19	11.0	8.9	12.9	10.5	1.0	16	10.7	7.7	12.9	10.7	1.3
Internostril distance	11	6.3	20.9	14.9	4.8	19	11.1	9.3	11.5	10.3	0.7	16	9.1	6.6	10.7	9.1	1.0
Posterior nostril-orbit distance	11	4.9	21.2	13.0	7.1	19	5.8	4.3	7.0	6.0	0.6	16	8.0	4.4	9.6	7.5	1.5
Head depth at supra occipital	11	37.7	57.0	49.4	6.6	19	49.9	48.0	51.5	49.5	1.1	16	38.2	35.7	41.6	38.9	1.7
Head depth at interorbital	6	23.7	38.2	32.1	5.6	19	38.1	34.5	43.5	37.5	1.9	16	28.0	26.3	29.9	28.2	1.1
Head width at posterior nostril	6	53.3	60.3	56.6	2.4	19	61.9	58.6	64.7	62.1	1.2	16	56.2	54.2	59.4	57.5	1.4
Dorsal head length	6	84.0	91.9	88.7	3.1	19	86.0	81.8	87.9	85.2	1.3	16	88.6	85.4	90.9	87.7	1.4