Taxonomic revision of the Quasipaa verrucospinosa complex (Amphibia, Dicroglossidae) in Vietnam, with descriptions of two new species Author Pham, Cuong The 0000-0001-5158-4526 Institute of Biology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, 10072 Hanoi, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, 10072 Hanoi, Vietnam Author Hoang, Chung Van 0000-0002-0709-974X Institute of Biology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, 10072 Hanoi, Vietnam Author Phan, Tien Quang 0000-0002-2738-5364 Institute of Biology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, 10072 Hanoi, Vietnam Author Pham, Anh Van 0000-0002-6023-3418 Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, 10072 Hanoi, Vietnam Author Ong, An Vinh 0000-0003-3683-3832 Faculty of Environmental Sciences, University of Science, Vietnam National University, Hanoi, 334 Nguyen Trai Road, 11400 Hanoi, Vietnam Author Nguyen, Vien Hong Thi 0000-0002-3744-3054 Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, 10072 Hanoi, Vietnam & Department of Zoology, Vinh University, 182 Le Duan Road. Vinh City, Nghe An Province, Vietnam Author Ziegler, Thomas 0000-0002-4797-609X Faculty of Resources and Environment, Thai Nguyen University of Sciences, Thai Nguyen University, Tan Thinh Ward, Thai Nguyen City, Thai Nguyen 250000, Vietnam & AG Zoologischer Garten Köln, Riehler Strasse 173, D- 50735 Cologne, Germany Author Nguyen, Truong Quang 0000-0002-6601-0880 Institute of Biology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, 10072 Hanoi, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, 10072 Hanoi, Vietnam text ZooKeys 2025 2025-06-05 1240 139 175 journal article 310261 10.3897/zookeys.1240.147337 ee5d54b2-c157-4a10-8d84-a74f452ec79c 562CF389-0F40-4FA2-8171-CC51E7AFA2A7

Table 5. Measurements (in mm) and proportions of Quasipaa verrucospinosa (M = Male, F = Female, SD = standard deviation; for other abbreviations see Material and methods).

	Quasipaa verrucospinosa				Quasipaa delacouri
Voucher	Min – Max (n = 7)	Mean ± SD (n = 7)	Min – Max (n = 7)	Mean ± SD (n = 7)	Min – Max (n = 3)	Mean ± SD (n = 3)	Min – Max (n = 3)	Mean ± SD (n = 3)
Sex	M	M	F	F	M	M	F	F
SVL	84.5–105.5	100.7 ± 6.6	75.2–94.7	86.8 ± 7.4	95.9–104.9	99.8 ± 4.6	90.5–98.4	94.1 ± 4.0
HL	31.1–39.9	38 ± 2.8	29.6–34.9	32.8 ± 20	33.5–35.6	34.3 ± 1.1	33.4–36.5	35.17 ± 1.6
HW	35.7–44.7	42.9 ± 3	33.3–39.6	37 ± 2.2	33.5–37.6	36.7 ± 1.0	34.6–40.0	37, 1 ± 2.7
MN	21.9–27.3	26 ± 2	21.6–25.4	23.7 ± 1.3	27.8–28.9	28.4 ± 0.6	23.9–29.5	27.2 ± 2.9
MFE	17.7–21.1	20.3 ± 1.1	16.6–20.1	18.6 ± 1.1	22.7–24.7	23.6 ± 1.0	17.8–25.7	22.4 ± 4.1
MBE	10.2–13.4	11.7 ± 1.1	8.6–11.8	10.5 ± 1	15.6–16.8	16.0 ± 0.7	14.5–17.4	15.7 ± 1.5
RL	13.2–15.4	15.1 ± 0.7	11.9–14.2	13.1 ± 1	13.2–14.7	14.1 ± 0.8	11.5–15.2	13.7 ± 1.9
ED	8.6–10.8	10.3 ± 0.7	9–11.5	10 ± 1	11.3–12.9	11.9 ± 0.8	11.1–12.9	12.3 ± 1.0
UEW	6.5–8.4	7.7 ± 0.6	6.0–7.5	7.0 ± 0.6	7.9–9.6	8.8 ± 0.9	7.1–8.8	7.8 ± 0.9
IND	7.3–9.7	8.7 ± 0.8	6.0–8.4	7.4 ± 0.9	10.2–11.2	10.6 ± 0.5	8.9–10.9	10.0 ± 1.0
IOD	6.9–8.9	8.2 ± 0.6	6.2–7.9	7.3 ± 0.6	6.2–7.3	6.8 ± 0.6	6.3–7.7	7.1 ± 0.7
DAE	12.1–14.6	13.5 ± 1	11.3–13.8	12.4 ± 0.9	15.5–6.3	15.9 ± 0.4	13.9–16.8	15.5 ± 1.5
DPE	22.8–28.2	26.3 ± 1.6	21.3–25.9	23.9 ± 1.6	24.5–24.9	24.7 ± 0.8	23.2–25.6	24.4 ± 1.2
NS	6.0–9.0	7.5 ± 0.9	6.1–8.1	7.0 ± 0.8	7.7–8.1	7.9 ± 0.2	6.5–8.8	7.7 ± 1.2
EN	5.7–7.4	6.8 ± 0.6	5.5–6.4	6.0 ± 0.3	5.8–6.6	6.3 ± 0.4	5.8–7.2	6.4 ± 0.7
TD	4.8–6.3	5.4 ± 0.5	4.6–5.9	5.0 ± 0.5	3.0–3.2	3.1 ± 0.1	2.8–3.1	2.9 ± 0.2
TYE	5.1–6.8	6.3 ± 0.6	4.6–6.6	5.6 ± 0.7	6.4–6.9	6.7 ± 0.3	5.5–5.8	5.6 ± 0.2
UAL	13.0–21.0	20.1 ± 2.8	13.4–16.7	15.5 ± 1.4	14.3–15.4	14.8 ± 0.5	14.5–16.2	15.4 ± 0.8
FAL	40.2–50.2	48.5 ± 3.5	34.2–41.9	38.4 ± 2.6	41.1–43.3	42.4 ± 1.2	40.4–44.5	41.8 ± 2.3
FeL	47.2–55.3	52.0 ± 3.1	38.4–49.3	44.8 ± 3.8	46.7–48.9	47.7 ± 1.1	46.9–49.8	49.1 ± 1.5
TbL	47.0–55.9	53.1 ± 3	40.4–48.9	45.7 ± 3.1	50.2–53.8	51.6 ± 1.9	50.0–54.7	51.5 ± 2.4
TbW	16.3–21.9	18.9 ± 1.8	12.8–17.3	15.0 ± 1.7	17.3–17.7	17.5 ± 0.2	14.9–17.2	15.8 ± 1.3
FoL	62.9–73.6	70.7 ± 3.5	54.7–65.5	60.8 ± 4.2	67.5–73.1	70.4 ± 2.8	68.5–71.3	69.7 ± 1.4
IMT	7.5–9.5	8.9 ± 0.7	5.4–8.1	6.9 ± 1	8.3–8.5	8.4 ± 0.1	7.8–9.0	8.5 ± 0.6
HL / SVL	0.37–0.39	0.38 ± 0.01	0.36–0.39	0.38 ± 0.01	0.32–0.36	0.34 ± 0.02	0.37–0.38	0.37 ± 0.01
HW / SVL	0.41–0.44	0.43 ± 0.01	0.41–0.44	0.43 ± 0.01	0.35–0.38	0.37 ± 0.02	0.38–0.41	0.39 ± 0.02
RL / SVL	0.14–0.16	0.15 ± 0.01	0.14–0.16	0.15 ± 0.01	0.13–0.15	0.14 ± 0.01	0.13–0.15	0.14 ± 0.01
HL / HW	0.85–0.91	0.88 ± 0.02	0.86–0.91	0.89 ± 0.02	0.92–0.95	0.94 ± 0, 02	0.91–0.97	0.95 ± 0.03
ED / RL	0.65–0.72	0.69 ± 0.03	0.65–0.85	0.77 ± 0.08	0.80–0.89	0.85 ± 0.05	0.85–0.97	0.90 ± 0.06
TYE / TD	0.94–1.34	1.16 ± 0.14	0.95–1.37	1.14 ± 0.16	2.00–2.16	2.15 ± 0.15	1.84–2.00	1.93 ± 0.08
TD / ED	0.46–0.60	0.53 ± 0.05	0.41–0.60	0.50 ± 0.06	0.23–0.27	0.26 ± 0.02	0.22–0.25	0.24 ± 0.01
TbL / SVL	0.51–0.56	0.53 ± 0.01	0.51–0.56	0.53 ± 0.02	0.48–0.55	0.55 ± 0, 04	0.55–0.57	0.56 ± 0.01
TbL / TbW	2.56–3.00	2.81 ± 0.15	2.78–3.34	3.06 ± 0.18	2.85–3.04	2.94 ± 0.09	3.18–3.48	3.34 ± 0.15