The contribution of environmental factors to fish assemblages in the Río Acahuapa, a small drainage in Central America Author Álvarez, Francisco S. Departamento de Investigación, UDP Ciencias Neotropicales, Calle Francisco Campos 166, Colonia Escalón, CP 1101, San Salvador, El Salvador samuel_biologo@hotmail.com Author Matamoros, Wilfredo A. Instituto de Ciencias Biológicas, Universidad de Ciencias y Artes de Chiapas, Libramiento Norte Poniente 1150, Colonia Lajas Maciel, CP 29039, Tuxtla Gutiérrez, Chiapas, México wilmatamoros@yahoo.com Author Chicas, Francisco A. Facultad de Ciencias Naturales y Matemática, Escuela de Biología, Universidad de El Salvador, Final Av. Estudiantes Héroes y Mártires del 30 de julio, Ciudad Universitaria, CP 1101, San Salvador, El Salvador francisco.chicas@ues.edu.sv text Neotropical Ichthyology 2017 2017-10-05 15 3 e 170023 1 11 http://dx.doi.org/10.1590/1982-0224-20170023 journal article 10.1590/1982-0224-20170023 1982-0224 20719581

Tab. 2. List of species classified according with their tolerance to salinity, abundance, standard length (maximum, minimum, average and standard deviation) and occurrence by elevation; High (352-519), Medium (185-351), and Low (19-184) meters above the sea level in the main rivers of the Río Acahuapa basin, San Vicente, El Salvador, 2011-2012. Shaded data represents the month of occurrence, the black circles represent occurrence by elevation. a Considered amphidromous or catadromous species by different authors based on direct or indirect evidence (Cruz 1987; Ribeiro & Villalobos 2010; Lorion et al. 2011; Smith & Kwak 2014b, 2014a). b Considered diadromous or amphidromous species by different authors based on direct or indirect evidence (Lorion et al. 2011; Smith & Kwak 2014b). c Considered diadromous or amphidromous species by different authors based on direct or indirect evidence (Lyons 2005; Smith & Kwak 2014b).

Family	Species	N	Classification tolerance an origin	Standard length (cm) minimum/ maximum	Average size/ standard deviation	Samples
						June			August			November			February			May
						H	M	L	H	M	L	H	M	L	H	M	L	H	M	L
Characidae	Astyanax aeneus (Günther, 1860) Roeboides bouchellei Fowler, 1923	439 8	Primary Primary	4.22-9.00 5.00-6.50	6.38±1.41 5.93±0.81	●	●	● ●	●	●	●	●	●	● ●	●	●	●	●	●	● ●
Heptapteridae	Rhamdia guatemalensis (Günther, 1864)	62	Primary	3.37-14.50	8.81±3.48		●	●		●				●		●
	Rhamdia laticauda (Kner, 1858)	65	Primary	3.50-19.72	11.14±4.26			●		●	●		●						●
Ariidae	Cathorops steindachneri (Gilbert & Starks, 1904)	18	Marine-estuarine	6.06-9.42	7.74±2.38			●												●
Mugilidae	Agonostomus monticola (Bancroft, 1834)	222	Diadromousa	4.64-25.50	7.95±1.49		●	●		●	●			●			●			●
	Mugil curema Valenciennes, 1836	8	Marine-estuarine	3.71-4.00	3.86±0.21			●			●
Anablepidae	Anableps dowei Gill, 1861	8	Secondary	7.00-16.50	10.41±2.43												●			●
	Poecilia gillii (Kner, 1863)	55	Secondary	4.00-10.25	6.77±1.81					●	●		●	●						●
	Poecilia marcellinoi Poeser, 1995	768	Secondary	3.00-6.80	4.24±0.94	●	●	●		●	●	●	●	●	●	●	●	●	●	●
Poeciliidae	Poecilia butleri Jordan, 1889	802	Secondary	2.92-5.50	3.56±1.31		●			●			●			●			●
	Poecilia salvatoris Regan, 1907	2721	Secondary	1.70-9.75	4.92±1.48	●	●	●	●	●	●	●	●	●	●	●	●	●	●	●
	Poeciliopsis pleurospilus (Günther, 1866)	741	Secondary	2.00-6.00	3.56±0.88	●	●	●	●	●	●	●	●	●	●	●	●	●	●	●
	Poeciliopsis turrubarensis (Meek, 1912)	107	Secondary	2.44-7.00	3.86±1.64						●			●			●			●
Synbranchidae	Synbranchus marmoratus Bloch, 1795	2	Secondary	25.00-40.00	32.50±10.61								●						●
	Centropomus medius Günther, 1864	1	Marine-estuarine	33.00	---												●
Centropomidae	Centropomus nigrescens Günther, 1864	5	Marine-estuarine	14.20	---															●
	Centropomus robalito Jordan & Gilbert 1882	1	Marine-estuarine	17.00	---												●
Sciaenidae	Bairdiella armata Gill, 1863	1	Marine-estuarine	9.00	---															●
	Amatitlania nigrofasciata (Günther, 1867)	446	Secondary	1.50-10.00	5.93±1.65	●	●	●	●	●	●	●	●	●	●	●	●	●	●	●
	Amphilophus longimanus (Günther, 1867)	1	Secondary	5.50	---			●
	Amphilophus trimaculatus (Günther, 1867)	1	Secondary	13.00	---															●
Cichlidae	Amphilophus macracanthus (Günther, 1864)	1	Secondary	23.00	---												●
	Oreochromis niloticus (Linneaus, 1758)	1	Secondary: Exotic	9.00	---															●
	Parachromis managuensis (Günther, 1867)	1	Secondary: Exotic	25.00	---									●
	Parachromis motaguensis (Günther, 1867)	8	Secondary	5.33-8.00	6.02±1.53		●			●	●
	Dormitator latifrons (Richardson, 1844)	36	Marine-estuarine	6.00-12.00	8.28±2.53			●						●			●			●
Eleotridae	Eleotris picta Kner, 1863	1	Marine-estuarine	37.00	---												●
	Gobiomorus maculatus (Günther, 1859)	23	Marine-estuarine	7.00-17.00	9.62±3.71			●			●			●			●			●
Gobiidae	Awaous banana (Valenciennes, 1837)	3	Diadromousb	9.00-13.10	10.05±1.48						●									●
	Sicydium multipunctatum Regan, 1906	183	Diadromousc	6.12-19.25	8.98±2.28			●			●		●	●		●	●			●
Achiridae	Trinectes fonsecensis (Günther, 1862)	1	Marine-estuarine	9.00	---															●
Paralichthyidae	Cyclopsetta panamensis (Steindachner, 1876)	2	Marine-estuarine	10.00	---															●
	Total	6,742			Total species	5	9	15	4	11	14	5	10	14	5	8	15	5	8	21
					Primary	1	2	4	1	3	2	1	2	3	1	2	1	1	2	2
					Secondary	4	6	5	3	7	7	4	7	7	4	5	7	4	6	9
					Marine-estuarine	0	0	4	0	0	2	0	0	2	0	0	5	0	0	7
					Diadromous	0	1	2	0	1	3	0	1	2	0	1	2	0	0	3