A new species of Dixonius Bauer, Good, and Branch, 1997 (Squamata: Gekkonidae) ranging through the Cardamon Mountains of Cambodia to the southern Mekong Delta of Vietnam Author Grismer, L. Lee 0000-0001-8422-3698 Department of Herpetology, San Diego Natural History Museum, P. O. Box 121390, San Diego, California 92112 USA grismer@lasierra.edu Author Neang, Thy Wild Earth Allies, 77 a, Street Beton, Bayap Village, Sangkat Phnom Penh Thmei, Khan Sen Sok, Phnom Penh, Cambodia Author Tri, Ngo Van Institute of Life Sciences, Vietnam Academy of Science and Technology, 9 / 621 Vo Nguyen Giap Highway, Linh Xuan Ward, Thu Đuc City, Hochiminh City, Vietnam Author Grismer, Jesse L. 0000-0001-8422-3698 grismer@lasierra.edu text Zootaxa 2025 2025-11-05 5717 2 180 206 https://doi.org/10.11646/zootaxa.5717.2.2 journal article 313845 10.11646/zootaxa.5717.2.2 fb2a4574-389f-4553-9564-f38dddd3a22f 1175-5326 17890208 D8B09AF0-D8BB-45B5-BDF6-9224EBFBA438

TABLE 5. Mean, one standard deviation, and range (in parentheses) of the size-corrected morphometric and meristic characters of Dixonius used in the analyses of this study.

species	SVL	HL	HW	HD	EL	ED	EN	ES	EE	IN
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aaronbaueri	1.55 ± 0.03 (1.5–1.59)	0.96 ± 0.01 (0.95–0.98)	0.75 ± 0.01 (0.74–0.76)	0.57 ± 0.01 (0.55–0.58)	-0.37 ± 0.03 (-0.4–-0.34)	0.29 ± 0.02 (0.28–0.3)	0.32 ± 0 (0.31–0.32)	0.48 ± 0.01 (0.46–0.49)	0.41 ± 0 (0.41–0.41)	-0.05 ± 0 (- 0.05–-0.05)
chotjuckdikuli	1.63 ± 0.03 (1.58–1.66)	1.09 ± 0.01 (1.07–1.11)	0.92 ± 0.02 (0.9–0.95)	0.73 ± 0.03 (0.68–0.76)	-0.27 ± 0.04 (-0.31–-0.22)	0.49 ± 0 (0.48–0.49)	0.54 ± 0.03 (0.49–0.57)	0.68 ± 0.02 (0.66–0.7)	0.57 ± 0.02 (0.54–0.6)	0.26 ± 0.04 (0.2–0.31)
dulayaphitakorum	1.62 ± 0.04 (1.58–1.66)	1.11 ± 0.01 (1.09–1.12)	0.92 ± 0.01 (0.91–0.94)	0.69 ± 0.03 (0.66–0.73)	0.06 ± 0.03 (0.03–0.1)	0.46 ± 0.03 (0.42–0.48)	0.53 ± 0.01 (0.52–0.54)	0.65 ± 0.02 (0.63–0.66)	0.55 ± 0.01 (0.54–0.56)	0.26 ± 0.01 (0.25–0.27)
fulbrighti	1.55 ± 0.08 (1.48–1.66)	1.03 ± 0.01 (1.02–1.04)	0.81 ± 0.04 (0.76–0.84)	0.54 ± 0.05 (0.49–0.61)	0.05 ± 0.07 (-0.03–0.13)	0.33 ± 0.02 (0.31–0.35)	0.45 ± 0.03 (0.41–0.48)	0.55 ± 0.09 (0.44–0.62)	0.49 ± 0.03 (0.44–0.52)	0.11 ± 0.01 (0.11–0.12)
gialaiensis	1.62 ± 0.06 (1.56–1.68)	1.07 ± 0 (1.06– 1.07)	0.89 ± 0 (0.89–0.89)	0.73 ± 0.01 (0.72–0.73)	0.04 ± 0 (0.04–0.05)	0.47 ± 0 (0.47–0.47)	0.5 ± 0.01 (0.49–0.51)	0.63 ± 0 (0.63–0.64)	0.51 ± 0 (0.51–0.52)	0.14 ± 0.02 (0.12–0.15)
kaweesaki	1.58 ± 0.06 (1.5–1.62)	1.06 ± 0 (1.06– 1.06)	0.87 ± 0.02 (0.84–0.89)	0.72 ± 0.01 (0.7–0.73)	-0.33 ± 0.06 (-0.41–-0.26)	0.52 ± 0 (0.52–0.53)	0.53 ± 0.03 (0.5–0.57)	0.64 ± 0.01 (0.63–0.65)	0.52 ± 0.01 (0.51–0.53)	0.06 ± 0.06 (-0.03–0.12)
lao	1.67 ± 0.1 (1.55–1.74)	1.11 ± 0.02 (1.09–1.13)	0.94 ± 0.01 (0.93–0.94)	0.71 ± 0.01 (0.7–0.72)	0.15 ± 0.02 (0.12–0.17)	0.52 ± 0.02 (0.51–0.54)	0.57 ± 0.04 (0.54–0.62)	0.69 ± 0.03 (0.67–0.72)	0.57 ± 0.01 (0.56–0.58)	0.18 ± 0.01 (0.17–0.2)
luuhoangorum sp. nov	1.63 ± 0.07 (1.48–1.7)	1.08 ± 0.05 (0.98–1.13)	0.88 ± 0.05 (0.78–0.94)	0.69 ± 0.06 (0.58–0.77)	0.15 ± 0.09 (-0.07–0.29)	0.4 ± 0.07 (0.26–0.49)	0.5 ± 0.07 (0.39–0.65)	0.65 ± 0.05 (0.55–0.72)	0.5 ± 0.09 (0.34–0.66)	0.18 ± 0.06 (0.06–0.3)
mekongensis	1.69 ± 0.02 (1.66–1.71)	1.18 ± 0.01 (1.17–1.19)	0.97 ± 0.05 (0.94–1.02)	0.71 ± 0.11 (0.64–0.84)	-0.05 ± 0 (- 0.05–-0.05)	0.52 ± 0.01 (0.52–0.53)	0.58 ± 0.03 (0.57–0.61)	0.72 ± 0.03 (0.7–0.75)	0.63 ± 0.05 (0.6–0.68)	0.31 ± 0.03 (0.29–0.34)
minhlei	1.65 ± 0.03 (1.61–1.68)	0.86 ± 0.01 (0.85–0.87)	0.82 ± 0.04 (0.79–0.89)	0.67 ± 0.02 (0.65–0.71)	0.12 ± 0.04 (0.08–0.18)	0.45 ± 0.02 (0.42–0.49)	0.55 ± 0.02 (0.52–0.57)	0.68 ± 0.03 (0.64–0.7)	0.56 ± 0.02 (0.53–0.59)	0.17 ± 0.02 (0.14–0.21)
muangfuangensis	1.69 ± 0.1 (1.58–1.75)	1.15 ± 0.02 (1.13–1.16)	0.98 ± 0.01 (0.97–0.99)	0.78 ± 0.01 (0.77–0.78)	0.21 ± 0.03 (0.18–0.25)	0.47 ± 0.03 (0.44–0.5)	0.54 ± 0 (0.54–0.54)	0.68 ± 0.01 (0.67–0.69)	0.66 ± 0 (0.65–0.66)	0.19 ± 0.01 (0.18–0.2)
noctivagus	1.66 ± 0.04 (1.55–1.73)	1.09 ± 0.03 (1.01–1.12)	0.93 ± 0.02 (0.9–0.95)	0.74 ± 0.02 (0.7–0.79)	0.16 ± 0.05 (0.04–0.23)	0.61 ± 0.03 (0.53–0.65)	0.54 ± 0.06 (0.49–0.69)	0.68 ± 0.01 (0.66–0.7)	0.53 ± 0.03 (0.48–0.57)	0.2 ± 0.04 (0.13–0.28)
pawangkhananti	1.62 ± 0.01 (1.61–1.63)	1.12 ± 0.01 (1.11–1.12)	0.9 ± 0 (0.89–0.9)	0.68 ± 0.01 (0.67–0.68)	-0.18 ± 0.04 (-0.22–-0.15)	0.47 ± 0.01 (0.47–0.48)	0.56 ± 0.02 (0.55–0.58)	0.69 ± 0.03 (0.66–0.71)	0.57 ± 0.02 (0.55–0.58)	0.18 ± 0 (0.18–0.18)
sambhupura	1.58 ± 0.14 (1.29–1.69)	1.09 ± 0.02 (1.05–1.11)	0.9 ± 0.02 (0.87–0.93)	0.74 ± 0.03 (0.71–0.81)	-0.1 ± 0.07 (- 0.29–-0.04)	0.43 ± 0.02 (0.39–0.47)	0.55 ± 0.02 (0.51–0.58)	0.68 ± 0.02 (0.66–0.7)	0.55 ± 0.02 (0.53–0.59)	0.16 ± 0.03 (0.11–0.2)
siamensis	1.58 ± 0.03 (1.56–1.6)	1.08 ± 0 (1.08– 1.08)	0.89 ± 0 (0.89–0.89)	0.72 ± 0 (0.72–0.72)	-0.13 ± 0 (- 0.13–-0.13)	0.41 ± 0 (0.41–0.41)	0.52 ± 0 (0.52–0.52)	0.66 ± 0 (0.66–0.66)	0.57 ± 0 (0.57–0.57)	0.17 ± 0 (0.17–0.17)
somchanhae	1.61 ± 0.05 (1.55–1.67)	1.06 ± 0.01 (1.05–1.08)	0.91 ± 0.01 (0.9–0.92)	0.7 ± 0.03 (0.66–0.74)	0.17 ± 0.04 (0.1–0.2)	0.47 ± 0.03 (0.43–0.51)	0.47 ± 0.03 (0.42–0.49)	0.64 ± 0.02 (0.62–0.68)	0.49 ± 0.02 (0.47–0.51)	0.16 ± 0.06 (0.1–0.24)
vietnamensis	1.56 ± 0.09 (1.41–1.66)	0.83 ± 0.02 (0.8–0.85)	0.82 ± 0.03 (0.77–0.86)	0.65 ± 0.05 (0.58–0.72)	-0.03 ± 0.05 (-0.13–0.05)	0.41 ± 0.03 (0.35–0.46)	0.47 ± 0.03 (0.42–0.51)	0.61 ± 0.04 (0.55–0.68)	0.48 ± 0.05 (0.36–0.55)	0.1 ± 0.07 (0.02–0.28)
species	IO	FA	TBL	AG	SL	IL	IOS	VS	T4L
aaronbaueri	0.49 ± 0.03 (0.46–0.52)	0.59 ± 0.01 (0.58–0.6)	0.73 ± 0.02 (0.7–0.75)	1.15 ± 0 (1.14–1.15)	8.38 ± 0.48 (8–9)	7 ± 0.41 (6.5–7.5)	9 ± 0.82 (8–10)	18 ± 0 (18– 18)	12.75 ± 1.26 (11–14)
chotjuckdikuli	0.23 ± 0.05 (0.17–0.31)	0.75 ± 0.02 (0.71–0.77)	0.84 ± 0.02 (0.82–0.88)	1.29 ± 0.02 (1.27–1.32)	7.75 ± 0.52 (7–9)	6.33 ± 0.61 (6–8)	7.67 ± 0.82 (6–8)	17.67 ± 0.82 (16–18)	16 ± 0.89 (15–17)
dulayaphitakorum	0.3 ± 0.01 (0.29–0.31)	0.68 ± 0.03 (0.63–0.72)	0.84 ± 0.01 (0.83–0.85)	1.27 ± 0.02 (1.25–1.3)	8 ± 0 (8–8)	7 ± 0 (7–7)	9.6 ± 0.55 (9–10)	22 ± 0 (22– 22)	13.6 ± 1.08 (12–15)
fulbrighti	0.21 ± 0.02 (0.19–0.22)	0.66 ± 0.03 (0.62–0.7)	0.75 ± 0.03 (0.72–0.8)	1.18 ± 0.02 (1.16–1.2)	8 ± 0.41 (7.5–9)	6.25 ± 0.65 (6–7)	8.25 ± 0.5 (8–9)	23 ± 0.82 (22–24)	14 ± 0.41 (13.5–14.5)
gialaiensis	0.11 ± 0.03 (0.08–0.13)	0.75 ± 0.03 (0.73–0.79)	0.83 ± 0.01 (0.82–0.84)	1.23 ± 0.02 (1.2–1.25)	7.5 ± 0.5 (7–8)	6.33 ± 0.58 (6–7)	7 ± 0 (7–7)	20.33 ± 1.15 (19–21)	14.17 ± 0.29 (14–14.5)
kaweesaki	0.59 ± 0.02 (0.56–0.61)	0.71 ± 0.02 (0.69–0.73)	0.84 ± 0.03 (0.81–0.87)	1.22 ± 0.02 (1.2–1.25)	10.12 ± 0.25 (10–11)	7.38 ± 0.63 (7–8)	6.5 ± 0.58 (6–7)	24 ± 0 (24– 24)	15 ± 0 (15–15)
lao	0.16 ± 0.05 (0.12–0.21)	0.79 ± 0.02 (0.78–0.81)	0.87 ± 0.01 (0.86–0.87)	1.29 ± 0 (1.28–1.29)	9 ± 0.5 (9–10)	7.83 ± 0.29 (7–8)	8.33 ± 0.58 (8–9)	23.33 ± 0.58 (23–24)	15 ± 0 (15–15)
luuhoangorum sp. nov	0.47 ± 0.06 (0.39–0.57)	0.77 ± 0.05 (0.65–0.83)	0.84 ± 0.06 (0.73–0.9)	1.27 ± 0.08 (1.09–1.33)	7.76 ± 0.66 (7–9)	6.26 ± 0.44 (6–7)	10.12 ± 1.17 (7–12)	17.71 ± 2.08 (14–22)	14.24 ± 1.19 (13–17)
mekongensis	0.34 ± 0 (0.34– 0.34)	0.78 ± 0.03 (0.77–0.82)	0.83 ± 0.02 (0.82–0.85)	1.32 ± 0.01 (1.32–1.33)	8.33 ± 0.29 (8–9)	6.5 ± 0.5 (6–7)	8.33 ± 0.58 (8–9)	22.67 ± 1.15 (22–24)	13.67 ± 1.53 (12–15)
minhlei	0.57 ± 0.03 (0.53–0.6)	0.8 ± 0.02 (0.78–0.83)	0.86 ± 0.02 (0.85–0.89)	1.33 ± 0.06 (1.28–1.44)	7.75 ± 0.42 (7–8)	6.42 ± 0.38 (6–7)	7.67 ± 1.21 (7–10)	21.33 ± 1.37 (20–23)	14.33 ± 1.03 (13–15)
muangfuangensis	0.33 ± 0.01 (0.32–0.34)	0.78 ± 0 (0.78– 0.78)	0.81 ± 0 (0.81–0.81)	1.35 ± 0.03 (1.31–1.38)	8.17 ± 0.29 (8–89)	6.5 ± 0.5 (6–7)	10 ± 0 (10–10)	22.33 ± 0.58 (22–23)	13.5 ± 1.32 (12–14.5)
noctivagus	0.25 ± 0.04 (0.19–0.35)	0.8 ± 0.03 (0.73–0.84)	0.86 ± 0.02 (0.83–0.9)	1.33 ± 0.02 (1.3–1.37)	7.94 ± 0.57 (7–9)	6.56 ± 0.51 (6–7)	10 ± 1.46 (8–13)	17.75 ± 1.65 (14–21)	11.69 ± 0.79 (11–13)
pawangkhananti	0.26 ± 0.02 (0.25–0.28)	0.76 ± 0.04 (0.71–0.79)	0.82 ± 0 (0.81–0.82)	1.25 ± 0 (1.25–1.25)	7.67 ± 0.58 (7–8)	6.33 ± 0.58 (6–7)	7 ± 0 (7–7)	16 ± 0 (16– 16)	14.67 ± 0.29 (14.5–15)
sambupura	0.57 ± 0.02 (0.53–0.59)	0.74 ± 0.02 (0.7–0.77)	0.83 ± 0.01 (0.81–0.85)	1.23 ± 0.04 (1.19–1.29)	7.9 ± 0.61 (7–9)	6.6 ± 0.52 (6–7)	9.7 ± 0.48 (9–10)	25.5 ± 1.08 (24–27)	12.7 ± 0.63 (12–14)
siamensis	0.48 ± 0 (0.48– 0.48)	0.7 ± 0 (0.7– 0.7)	0.82 ± 0 (0.82–0.82)	1.17 ± 0 (1.17–1.17)	8 ± 0 (8–8)	7 ± 0 (7–7)	10 ± 0 (10–10)	22.5 ± 3.54 (20–25)	14.5 ± 0.71 (14–15)
somchanhae	0.17 ± 0.04 (0.13–0.22)	0.71 ± 0.03 (0.68–0.76)	0.82 ± 0.02 (0.8–0.84)	1.27 ± 0.03 (1.24–1.31)	7.8 ± 0.45 (7–8)	5.6 ± 0.42 (5–6)	7.8 ± 0.45 (7–8)	23.8 ± 1.3 (23–26)	14.4 ± 0.89 (13–15)
vietnamensis	0.29 ± 0.16 (0.12–0.53)	0.67 ± 0.06 (0.55–0.76)	0.78 ± 0.03 (0.73–0.85)	1.2 ± 0.05 (1.12–1.27)	6.75 ± 1.18 (5–8)	6.29 ± 0.45 (6–7)	8.67 ± 0.98 (7–10)	19.25 ± 1.54 (15–21)	13.25 ± 0.4 (12.5–14)