Two New Species of Snailfishes (Cottiformes: Liparidae) from the Aleutian Islands, Alaska, and a Redescription of the Closely Related Careproctus candidus Author Gardner, Jennifer R. Author Orr, James W. Author Tornabene, Luke text Ichthyology & Herpetology 2023 2023-02-10 111 1 54 71 http://dx.doi.org/10.1643/i2022009 journal article 252075 10.1643/i2022009 3bd01deb-4887-4668-bfeb-1a5baf431a15 2766-1520 7847025

Table 1. Proportional morphometric and meristic data for Careproctus candidus and two new liparid species: Careproctus klisi and Careproctus bromius. Proportions in % standard length.

	Careproctus candidus				Careproctus klisi			Careproctus bromius
	n	Range	Mean6SD	n	Range	Mean6SD	n	Range	Mean6SD
Standard length	61	27.0–94.7		13	72.4–106.0		12	27.0–46.9
Head length	60	25.74–31.02	28.8 6 1.3	13	27.51–29.44	28.7 6 0.5	9	25.39–28.28	27.0 6 0.8
Head width	60	9.73–19.54	13.7 6 2.3	13	11.31–15.39	13.1 6 1.2	9	14.29–22.76	18.2 6 3.0
Body depth at anal-fin origin	60	16.88–25.87	20.5 6 2.0	13	21.41–28.96	25.0 6 2.2	9	12.79–20.25	16.9 6 2.4
Dorsal-fin origin to anal-fin origin length	59	23.68–32.70	27.5 6 2.0	13	26.49–33.71	30.2 6 2.0	9	22.47–29.64	25.6 6 2.8
Body depth at pelvic disc	60	20.16–28.36	24.7 6 2.0	13	23.11–28.87	26.5 6 1.8	9	14.93–26.17	21.9 6 3.8
Body depth at dorsal-fin origin	59	21.24–31.63	26.8 6 2.3	13	25.93–32.75	29.3 6 2.1	8	21.37–28.31	24.8 6 2.4
Body depth at anal-fin midpoint	59	6.94–12.30	8.9 6 1.2	13	10.22–15.38	12.1 6 1.5	9	6.90–8.14	7.4 6 0.4
Snout length	60	5.78–11.35	9.0 6 1.1	13	8.36–10.51	9.3 6 0.7	9	7.47–9.38	8.2 6 0.6
Orbit length	60	8.73–12.97	11.0 6 0.8	13	8.90–11.46	9.9 6 0.9	9	8.57–12.34	10.1 6 1.3
Bony interorbital width	57	1.68–5.12	3.2 6 0.8	12	3.01–4.23	3.4 6 0.3	9	3.38–5.93	4.7 6 0.9
Greatest interorbital width	58	3.51–11.36	7.4 6 1.8	12	6.88–9.12	7.9 6 0.8	9	6.40–12.34	10.0 6 2.0
Suborbital to maxilla length	60	1.04–5.27	2.9 6 0.8	13	2.21–4.63	3.3 6 0.7	9	1.42–5.18	3.2 6 1.3
Suborbital to mandible length	58	3.78–7.19	5.5 6 0.8	13	4.75–7.17	5.6 6 0.7	9	4.26–6.75	5.7 6 0.7
Mouth width	60	7.87–14.55	10.8 6 1.6	13	8.80–11.51	10.2 6 0.7	9	9.87–18.85	15.2 6 3.3
Maxilla length	60	9.37–15.36	11.6 6 1.2	13	9.48–11.48	10.7 6 0.5	9	11.81–17.86	14.7 6 1.8
Mandible length	60	7.80–15.70	11.1 6 1.8	13	9.91–12.74	11.4 6 0.9	9	12.07–19.31	14.8 6 1.9
Gill-slit length	60	3.93–7.40	5.5 6 0.8	13	4.15–7.50	5.5 6 1.0	9	3.85–6.17	5.4 6 0.8
Pectoral-fin length, upper lobe	60	17.45–24.07	20.6 6 1.4	13	17.23–22.54	20.3 6 1.7	9	14.96–21.38	18.4 6 2.3
Pectoral-fin length, lower lobe	60	11.76–22.21	17.9 6 2.3	13	14.11–23.08	17.7 6 2.7	9	10.62–19.51	15.8 6 3.0
Pectoral-fin length, notch ray length	59	4.32–10.59	7.7 6 1.5	13	6.13–10.37	8.4 6 1.3	9	6.74–9.87	8.3 6 1.2
Pre-dorsal-fin length	60	24.25–32.46	27.5 6 1.9	13	24.81–28.47	26.8 6 0.9	9	26.94–31.45	29.2 6 1.6
Pre-anal-fin length	60	34.69–46.90	40.5 6 3.3	13	37.99–45.99	40.6 6 2.3	9	44.09–49.87	46.7 6 2.1
Snout to pelvic-disc length	60	11.78–20.16	15.4 6 1.9	13	12.60–19.89	15.9 6 2.1	9	10.37–17.66	14.7 6 2.3
Snout to anus length	60	21.91–33.21	27.7 6 2.5	13	22.89–28.59	26.2 6 1.6	9	24.20–32.48	28.7 6 2.9
Pelvic-disc length	60	7.58–11.34	9.4 6 1.0	13	8.12–10.52	9.5 6 0.7	9	8.83–11.11	10.0 6 0.7
Pelvic-disc width	60	6.61–11.25	8.5 6 1.0	13	7.22–10.51	8.9 6 0.9	9	8.31–10.62	9.1 6 0.7
Pelvic-disc to anus length	60	1.23–6.00	3.1 6 1.2	13	1.33–4.27	2.5 6 1.0	9	1.43–5.45	3.7 6 1.6
Anus to anal-fin length	60	9.39–21.07	13.4 6 2.5	13	11.83–16.62	14.3 6 1.2	9	15.10–23.12	19.2 6 2.8
Caudal-fin length	54	11.91–16.18	14.3 6 1.0	13	13.49–14.88	14.1 6 0.5	9	11.94–15.24	13.5 6 1.1
Dorsal-fin attachment to caudal-fin length	57	1.95–5.62	3.4 6 0.7	12	2.89–4.91	4.2 6 0.6	9	1.98–3.91	2.7 6 0.6
Anal-fin attachment to caudal-fin length	57	2.85–7.19	5.1 6 1.0	13	5.12–7.73	6.1 6 0.7	9	3.13–6.19	4.4 6 1.1
Caudal-fin-base depth	59	1.41–2.57	1.9 6 0.3	13	1.68–2.51	2.1 6 0.2	9	1.58–2.86	2.2 6 0.5
Nostril-tube length	56	0.00–1.41	0.8 6 0.3	13	0.34–1.25	0.9 6 0.2	9	0.43–1.23	0.9 6 0.3
Urogenital-papilla length	55	0.00–1.81	0.7 6 0.5	13	0.00–2.90	0.7 6 0.8	9	0.00–0.57	0.2 6 0.2
Pyloric-caeca length	14	5.52–10.62	8.5 6 1.4	5	5.53–8.98	7.6 6 1.3	3	5.96–7.41	6.5 6 0.8
Pyloric caeca	11	16–24	18.5 6 2.0	4	17–21	19.2 6 2.1	3	13–16	15.0 6 1.7
Dorsal-fin rays	58	37–49	44.8 6 2.0	13	43–48	46.2 6 1.3	10	38–42	40.7 6 1.3
Anal-fin rays	56	35–41	38.4 6 1.5	13	37–41	39.1 6 1.3	12	32–37	34.2 6 1.4
Pectoral-fin rays	56	31–41	36.7 6 2.0	13	36–39	37.5 6 1.1	9	30–36	34.0 6 1.7
Rays of lower lobe of pectoral fin	58	6–12	8.0 6 1.0	13	7–9	8.4 6 0.7	9	7–8	7.1 6 0.3
Caudal-fin rays	46	10–11	10.8 6 0.4	10	10–11	10.5 6 0.5	10	11	11.0 6 0.0
Precaudal vertebrae	60	9–10	9.7 6 0.5	13	9–10	9.2 6 0.4	12	9–10	9.9 6 0.3
Caudal vertebrae	60	36–42	39.0 6 1.4	13	37–42	40.2 6 1.3	12	34–37	35.8 6 0.9
Total vertebrae	60	46–51	48.8 6 1.4	13	47–51	49.5 6 1.1	12	44–47	45.7 6 1.0
Tooth counts per row	30	4–10	6.9 6 0.9	13	5–10	7.0 6 1.2	8	4–12	7.5 6 1.5
Tooth rows	30	8–12	9.6 6 1.0	13	8–13	9.6 6 1.3	8	8–10	9.1 6 0.6
Gill rakers	35	6–10	8.0 6 1.0	6	7–10	8.3 6 1.5	8	6–8	6.6 6 0.7