Revision of Alpheus euphrosyne De Man, 1897 and A. microrhynchus De Man, 1897, with description of three new species and taxonomic remarks on several other morphologically and ecologically similar snapping shrimps (Malacostraca: Decapoda: Alpheidae) Author Anker, Arthur text Zootaxa 2023 2023-05-09 5282 1 1 115 http://dx.doi.org/10.11646/zootaxa.5282.1.1 journal article 54693 10.11646/zootaxa.5282.1.1 7912c179-a542-47d7-9b44-6af1c9185a23 1175-5326 7912292 DF418763-8F0E-44DD-97C4-B123A81A8DB4

TABLE 1. Comparison of main taxonomically useful characters (morphology, colour pattern, larval development, ecology) in the Alpheus euphrosyne De Man, 1897 —A. micro- rhynchus De Man, 1897 complex. Abbreviations used: A1—antennule / antennular; A2—antenna / antennal; P3, P4—third and fourth pereiopods, respectively.

Characters	Rostrum	Rostral carina	Rostro-orbital	Antennule, ventromesial carina
TABLE 1. Comparison of main taxonomically useful characters (morphology, colour pattern, larval development, ecology) in the Alpheus euphrosyne De Man, 1897 —A. micro- rhynchus De Man, 1897 complex. Abbreviations used: A1—antennule / antennular; A2—antenna / antennal; P3, P4—third and fourth pereiopods, respectively.
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Species			furrows	on peduncle article 1
A. euphrosyne	moderately developed, longer than broad, usually	usually present, very faint, restricted	barely distinct, very	with sharp, anteriorly directed tooth
	reaching half-length of 1st article of A1 peduncle	to rostrum, sometimes barely	shallow, sometimes
		distinct or absent	indistinct
A. eurydactylus	well developed, longer than broad, usually reaching	present, broad, blunt, flattening	distinct, shallow	broadly triangular, usually
	half-length of 1st article of A1 peduncle (not reaching	posterior to eye level		without tooth (with tooth in A. cf.
	0.3 length in A. cf. eurydactylus, ZRC 1992.8064)			eurydactylus, ZRC 1992.8064)
A. richardsoni	well developed, longer than broad, usually	present, broad, blunt, flattening	distinct, shallow	broadly triangular, usually with
	overreaching half-length of 1st article of A1 peduncle	posterior to eye level		small tooth
A. microrhynchus	very short, broader than long, not reaching 0.2 length	absent or barely distinct, flattening	indistinct	triangular, usually with small tooth
	of 1st article of A1 peduncle	between eyes
A. cyanoteles	short, as long as broad, not reaching 0.3 length of 1st	present, faint, blunt, restricted to	very shallow	triangular, usually with small tooth
	article of A1 peduncle	rostrum
A. nomurai sp. nov.	well developed, longer than broad, usually reaching	present, broad, blunt, flattening	rather distinct,	with sharp, anteriorly directed tooth
	half-length of 1st article of A1 peduncle	posterior to eye level	shallow
A. takla sp. nov.	short, longer than broad, usually reaching 0.3 length of	present, faint, blunt, flattening	usually distinct, very	broadly triangular, usually without
	1st article of A1 peduncle	between eyes	shallow	tooth
A. mangalis sp. nov.	well developed, longer than broad, usually reaching	present, faint, blunt, flattening	very shallow	broadly triangular, without tooth
	half-length of 1st article of A1 peduncle	between eyes
A. songkla	well developed, longer than broad, reaching half-length	present, faint, blunt, flattening	very shallow, barely	unknown (not examined in the type
	of 1st article of A1 peduncle	between eyes	distinct	specimen, with small tooth in some
				specimens of A. cf. songkla)
A. paludicola	very short, broader than long, not reaching 0.2 length	absent	indistinct	unknown (not examined)
	of 1st article of A1 peduncle
A. nipa	short, longer than broad, not reaching 0.3 length of 1st	present, moderate, not extending	very shallow or	triangular, without tooth
	article of A1 peduncle, with long erect setae	posterior to eye level	indistinct
A. bunburius	well developed, longer than broad, reaching half-length	present, broad, blunt, flattening	distinct, shallow	unknown (not examined)
	of 1st article of A1 peduncle	between eyes
A. pontederiae	well developed, longer than broad, reaching half-length	present, well-developed, flattening	distinct, shallow	with sharp, anteriorly directed tooth
	of 1st article of A1 peduncle	posterior to eye level
A. firmus	short, longer than broad, not reaching 0.3 length of 1st	present, narrow, blunt, flattening	distinct, shallow	triangular, without tooth
	article of A1 peduncle	posterior to eye level