Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae) Author De Mattia, Willy https://orcid.org/0000-0002-0056-467X Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria willy.demattia@icgeb.org Author Reier, Susanne Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria Author Haring, Elisabeth Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria text ZooKeys 2021 2021-12-14 1077 1 175 http://dx.doi.org/10.3897/zookeys.1077.67081 journal article http://dx.doi.org/10.3897/zookeys.1077.67081 1313-2970-1077-1 C28AD65A76F242CFBED7DFB3702CABCE 734088641608531C8E2CC69397B000ED

Table 4. Siciliaria / Sicania taxa examined (anatomy and/or molecular genetic data) with description of the variable penis-epiphallus transition: epiphallar formula. DNA = molecular genetic data available (Yes/No).

Taxon	Locality	DNA	Epiphallar formula
Siciliaria grohmanniana grohmanniana (Rossmaessler, 1836)	Italy, Sicily, Palermo, Monte Pellegrino, N side of the top plateau, 380 m asl, 38°10'55.26"N, 13°21'1.66"E, leg. W. De Mattia and J. Macor leg.	Y	1ER(PP+ELP)
	Italy, Sicily, Palermo, Monte Pellegrino, Santuario Santa Rosalia, 420 m asl, 38°10'4.41"N, 13°21'2.60"E, leg. W. De Mattia and J. Macor leg.	Y	1ER(PP+ELP)
Siciliaria grohmanniana addaurae ssp. nov. De Mattia, Reier & Haring	Italy, Sicily, Palermo, N side of Monte Pellegrino, Punta Priola, Grotte dell'Addaura, 115 m asl, 38°11'13.01"N, 13°21'6.91"E, W. De Mattia and J. Macor leg.	Y	1ER(PP)+ELP
Siciliaria septemplicata (Philippi, 1836)	Italy, Sicily, Palermo, N side of Monte Gallo near Sferracavallo, 40 m asl, 38°12'39.27"N, 13°17'21.26"E, W. De Mattia and J. Macor leg.	Y	1ER+2ER(PP+ELP)
	Italy, Sicily, Palermo, E side of Monte Gallo, 140 m asl, 38°13'5.61"N, 13°18'4.76"E, W. De Mattia and J. Macor leg.	Y	1ER+2ER(PP+ELP)
Siciliaria leucophryna (L. Pfeiffer, 1862)	Italy, Sicily, Palermo, Grotta Conza, 150 m asl, 38°11'13.61"N, 13°16'44.68"E, W. De Mattia and J. Macor leg.	Y	1ER+2ER(PP+ELP)
	Italy, Sicily, Palermo, Sferracavallo, via Plauto, 50 m asl, 38°11'13.01"N, 13°21'6.91"E, W. De Mattia and J. Macor leg.	Y	1ER(PP)+ELP
Siciliaria calcarae calcarae (Philippi, 1844)	Italy, Sicily, Alcamo, Monte Bonifato, top of the mountain, 640 m asl, 37°57'29.40"N, 12°57'33.64"E, A. Margelli leg.	N	1ER(PP)+ELP
	Italy, Sicily, Alcamo, Monte Bonifato, top of the mountain, 640 m asl, 37°57'29.40"N, 12°57'33.64"E, I. Niero leg.	Y	1ER(PP+ELP)
	Italy, Sicily, Alcamo, Monte Bonifato, west side of the mountain, over the quarry, 550 m asl, 37°57'16.92"N, 12°58'9.06"E, W. De Mattia and J. Macor leg.	N	1ER(PP+ELP)
	Italy, Sicily, Piana degli Albanesi, 500 m south of Portella Ginestra, northern cliffs of Monte Kumeta, 970 m asl, 37°58'13.35"N, 13°15'22.06"E, W. De Mattia and J. Macor leg.	N	1ER(PP)+2ER(ELP)
	Italy, Sicily, Castellammare del Golfo, Castello di Baida, W of the town along the road to Visicari, 300 m asl, 38° 02'41.64"N, 12°48'14.34"E, W. De Mattia and J. Macor leg.	N	1ER(PP)+2ER(ELP)
	Italy, Sicily, San Vito lo Capo, Castelluzzo, west cliffs E of town, 120 m asl, 38°06'25.71"N, 12°44'33.37"E, W. De Mattia and J. Macor leg.	Y	1ER(PP)+ELP
	Italy, Sicily, Castellammare del Golfo, Visicari, 395 m asl, 38°02'54.18"N, 12°48'26.61"E, W. De Mattia and J. Macor leg.	Y	1ER(PP+ELP)
Siciliaria calcarae calcarae (Philippi, 1844) (= Siciliaria adelina Kuester, 1847)	Italy, Sicily, Calatafimi, Castello Eufemio, 395 m asl, 37°57'45.67"N, 12°51'21.13"E, W. De Mattia and J. Macor leg.	N	1ER+2ER(PP)+ELP
Siciliaria calcarae belliemii (Brandt, 1961)	Italy, Sicily, Partinico, W side of the Mount Belliemi, 440 m asl, 38°00'22.47"N, 13°06'37.79"E, W. De Mattia and J. Macor leg.	Y	1ER+2ER(PP)+ELP
	Italy, Sicily, Alcamo, E side of the Calatubo Castle, 75 m asl, 38°00'54.80"N, 12°59'13.35"E, W. De Mattia and J. Macor leg.	Y	1ER+2ER(PP+ELP)
Siciliaria calcarae borgettensis ssp. nov. De Mattia, Reier & Haring	Italy, Sicily, Municipality of Borghetto, road to Romitello, 400 m asl, 38°02'59.53"N, 13°08'58.55"E, W. De Mattia and J. Macor leg.	Y	1ER(PP)+ELP
	Italy, Sicily, San Giuseppe Jato, quarry E of the town, 630 m asl, 37°58'15.07"N, 13°12'2.23"E, W. De Mattia and J. Macor leg.	Y	1ER+2ER(PP+ELP)
Siciliaria calcarae parajatinensis ssp. nov. De Mattia, Reier & Haring	Italy, Sicily, Monreale, W part of Monte Kumeta toward Jato Antica, 630 m asl, 37°57'8.18"N, 13°13'14.57"E, W. De Mattia and J. Macor leg.,	Y	1ER+2ER(PP+ELP)
Siciliaria calcarae orlandoi Liberto, Reitano, Giglio, Colomba & Sparacio, 2016	Italy, Sicily, Monreale, Bosco Ficuzza, Ponte Arcere, 470 m asl, 37°55'46.42"N, 13°23'5.22"E, W. De Mattia and J. Macor leg.	Y	1ER+2ER(PP+ELP)
Siciliaria calcarae cruenta ssp. nov. De Mattia, Reier & Haring	Italy, Sicily, Monreale, N side of Monte Gibilmesi, 890 m asl, 38°03'37.03"N, 13°12'38.53"E, W. De Mattia and J. Macor leg.	Y	1ER+2ER(PP)+ELP
Siciliaria ferrox Brandt, 1961	Italy, Sicily, Trabia, Contrada Sant'Onofrio, 170 m asl, 37°59'17.62"N, 13°36'51.11"E, W. De Mattia and J. Macor leg.	Y	?ER+PP(ELP)
Siliciaria tiberi tiberi (A. Schmidt, 1868)	Italy, Sicily, Terrasini, Capo Rama, 30 m asl, 38°08'19.06"N, 13°03'14.01"E, W. De Mattia and J. Macor leg.	Y	ER+PP(ELP)
Siciliaria tiberii alcamoensis Brandt, 1961 comb. nov.	Italy, Sicily, Cinisi, Piano Margi, 670 m asl, 38°08'58.56"N, 13°09'25.83"E, W. De Mattia and J. Macor leg.	Y	1ER(PP+ELP)
Siciliaria tiberii armettensis ssp. nov. De Mattia, Reier and Haring	Italy, Sicily, Carini, Grotta dei Puntali o dell'Armetta, 90 m asl, 38°08'58.56"N, 13°09'25.83"E, W. De Mattia and J. Macor leg.	Y	1ER+2ER+3ER(PP+ELP)
Siciliaria tiberii scalettensis Beckmann, 2004	Italy, Sicily, Cinisi, Portella Scaletta [locus typicus], 90 m asl, 38°10'35.53"N, 13°07'27.22"E, W. De Mattia and J. Macor leg.	Y	1ER+PP(ELP)
	Italy, Sicily, Cinisi, along the SS113 road from Villagrazia di Carini to Cinisi, 70 m asl, 38°10'42.45"N, 13°07'34.50"E, W. De Mattia and J. Macor leg.	Y	1ER+PP(ELP)
Sicania crassicostata (Pfeiffer, 1856)	Italy, Sicily, Castelluzzo, cliffs W of the Tonnara di Monte Cofano, 50 m asl, 38°6'44.89"N, 12°40'34.27"E, W. De Mattia and J. Macor leg.	Y	1ER+PP(ELP)
	Italy, Sicily, Custonaci, cliffs N of the Mangiapane cave, 63 m asl, 38°05'40.91"N, 12°40'10.98"E, W. De Mattia and J. Macor leg.	Y	1ER(PP)+ELP
Sicania eminens (A. Schmidt, 1868), comb. nov.	Italy, Sicily, Custonaci, Baglio Messina, 340 m asl, 38° 4'06.38"N, 12°41'48.68"E, W. De Mattia and J. Macor leg.,	Y	1ER+2ER(PP+ELP)
	Italy, Sicily, Custonaci, Buffara, NE side of the hollow, 150 m asl, 38°04'0.52"N, 12°41'1.34"E, W. De Mattia and J. Macor leg.	Y	1ER(PP+ELP)
	Italy, Sicily, Custonaci, contrada Scurati, crossing with Strada Provinciale 18, 65 m asl, 38°04'56.04"N, 12°40'9.86"E, W. De Mattia and J. Macor leg.	Y	1ER(PP)+2ER(ELP)
	Italy, Sicily, Custonaci, contrada Scurati, 60 m asl, 38° 05'18.10"N, 12°40'17.18"E, W. De Mattia and J. Macor leg.	Y	1ER(PP)+2ER(ELP)
	Italy, Sicily, Custonaci, contrada Scurati, limestone cliffs S of the village, 60 m asl, 38°06'44.89"N, 12°40'34.27"E, W. De Mattia and J. Macor leg.	Y	1ER(PP)+2ER(ELP)
Sicania nobilis nobilis (L. Pfeiffer, 1848), comb. nov.	Italy, Sicily, Castelluzzo, Monte Cofano E of Tonnara di Monte Cofano, 80 m asl, 38°06'22.04"N, 12°40'59.77"E, W. De Mattia and J. Macor leg.	N	1ER+2ER+3ER(PP+ELP)
	Italy, Sicily, San Vito lo Capo, boulders W of the town, 50 m asl, 38°10'51.85"N12°43'37.70"E, W. De Mattia and J. Macor leg.	Y	1ER+2ER+3ER(PP+ELP)
	Italy, Sicily, San Vito lo Capo, Castelluzzo, west cliffs E of town, 120 m asl, 38°06'25.71"N, 12°44'33.37"E, W. De Mattia and J. Macor leg.	Y	1ER+2ER(PP)+ELP
	Italy, Sicily, San Vito lo Capo, cliffs S of "El-Bahira" camping, 60 m asl, 38°08'48.81"N12°44'06.61"E, W. De Mattia and J. Macor leg.	Y	1ER+2ER(PP)+ELP
Sicania nobilis nobilis (L. Pfeiffer, 1848), comb. nov. (= Sicania episoma Brandt, 1961)	Italy, Sicily, San Vito lo Capo, Torre delle Usciere, 5 m asl, 38°10'30.07"N, 12°46'14.09"E, W. De Mattia and J. Macor leg.	Y	1ER+2ER+3ER(PP+ELP)
Sicania nobilis spezialensis (Nordsieck 1984), stat. nov., comb. nov.	Italy, Sicily, San Vito Lo Capo, Macari, E side of Monte Speziale, 75 m asl, 38°07'41.78"N, 12°44'09.24"E, W. De Mattia and J. Macor leg.	Y	1ER+2ER(PP+ELP)