Descriptions of two new shell-dwelling species of Metriaclima (Cichlidae) from Lake Malaŵi, Africa Author Miller, Torin A. Penn State University, 432 Forest Resources Building, University Park, Pennsylvania 16802. Author Konings, Adrianus F. Cichlid Press, PO Box 13608, El Paso, TX 79913. Author Stauffer, Jay R. Penn State University, 432 Forest Resources Building, University Park, Pennsylvania 16802. & Honorary Research Associate, South African Institute for Aquatic Biodiversity, Makhanda, RSA. text Zootaxa 2021 2021-10-15 5052 3 419 432 journal article 3996 10.11646/zootaxa.5052.3.7 af9fa8a9-6374-4b14-b037-fe6d44505dc1 1175-5326 5572218 B602FEFE-70FB-493E-BB25-3995F8F3A286

TABLE 1. Morphological and meristic data for Metriaclima ngarae from Ngara, Lake Malaŵi, PSU 13365, 13366, n=14. Ranges include holotype.

Variable	Holotype	Mean	Std Dev	Range
Standard length (mm)	59.1	56.6	3.7	49.6–62.4
Head length (mm)	19.1	18.7	1.2	17.0–21.4
Percent head length (%)
Snout length	34.6	34.9	1.2	30.8–36.4
Postorbital head length	39.5	40.8	1.7	36.8–42.7
Horizontal eye diameter	30.6	30.1	1.6	26.7–32.3
Vertical eye diameter	28.9	28.0	1.5	24.4–29.9
Preorbital depth	18.7	19.3	1.1	17.9–21.2
Cheek depth	35.3	34.6	2.5	30.3–38.6
Lower jaw length	25.0	26.0	2.8	20.7–30.4
Head depth	109.9	104.1	4.6	98.5–113.0
Inter-orbital width	34.4	33.4	2.1	30.7–37.4
Premaxillary pedicel length	15.0	12.2	2.9	7.0–15.8
Percent standard length (%)
Head length	32.3	33.1	0.9	32.0–34.5
Body depth	39.1	38.7	0.8	37.1–40.1
Snout to dorsal fin origin	34.9	35.7	1.3	33.9–37.7
Snout to pelvic fin origin	38.4	37.5	1.1	36.3–39.9
Dorsal fin base length	59.8	58.3	1.4	56.0–60.2
Anterior dorsal to anterior anal	50.1	50.1	0.8	48.8–51.5
Anterior dorsal to posterior anal	63.5	62.9	1.3	60.8–64.6
Posterior dorsal to anterior anal	36.6	36.0	1.2	34.2–37.7
Posterior dorsal to posterior anal	19.3	19.4	0.9	18.1–20.3
Posterior dorsal to ventral caudal	21.1	20.6	0.9	19.3–22.3
Posterior anal to dorsal caudal	25.1	23.2	1.1	20.6–25.1
Anterior dorsal to pelvic-fin origin	39.4	38.5	0.9	37.5–40.2
Posterior dorsal to pelvic-fin origin	65.2	61.5	1.4	58.7–65.2
Caudal peduncle length	14.9	14.0	1.0	11.8–15.4
Least caudal peduncle depth	16.3	16.2	0.6	15.5–17.3
Meristics		Mode	Frequency (%)	Range
Dorsal-fin spines	17	17	71.4	16–17
Dorsal-fin rays	8	8	71.4	7–8
Anal-fin spines	3	3	100.0	3–3
Anal-fin rays	8	8	100.0	8–8
Pectoral-fin rays	12	14	85.7	12–15
Pelvic-fin rays	7	7	85.7	6–7
Lateral line scales	32	31	57.1	30–32
Pored scales caudal	2	2	50.0	0–2
Cheek scale rows	4	4	100.0	4–4
Gill rakers 1st ceratobranchial	10	11	57.1	9–12
Gill rakers 1st epibranchial	5	5	50.0	4–5
Teeth outer left lower jaw	8	7	35.7	4–10
Tooth rows upper jaw	3	3	92.9	2–3
Tooth rows lower jaw	3	3	64.3	2–3
Standard length (mm)	54.4	45.8	5.0	36.4–54.4
Head length (mm)	17.7	15.1	1.4	12.4–17.7
Percent head length (%)
Snout length	28.3	31.0	2.3	26.2–35.0
Postorbital head length	41.9	42.7	1.8	39.0–46.0
Horizontal eye diameter	32.3	31.1	1.6	28.5–34.2
Vertical eye diameter	30.8	29.0	1.8	25.1–32.4
Preorbital depth	17.9	16.3	1.5	14.1–18.9
Cheek depth	31.1	26.3	3.6	21.1–33.5
Lower jaw length	26.9	26.0	3.0	21.6–33.5
Head depth	104.8	100.1	5.4	91.3–109.7
Inter-orbital width	26.2	27.3	1.5	23.6–29.7
Premaxillary pedicel length	15.7	13.7	1.7	10.9–16.4
Percent standard length (%)
Head length	32.7	33.0	0.8	31.5–34.4
Body depth	43.3	38.6	1.8	34.9–43.3
Snout to dorsal fin origin	39.5	35.7	1.3	32.6–39.5
Snout to pelvic fin origin	38.9	40.6	1.5	37.4–42.5
Dorsal fin base length	60.2	57.2	2.2	52.4–60.2
Anterior dorsal to anterior anal	55.3	51.8	1.7	48.6–55.3
Anterior dorsal to posterior anal	65.3	62.5	1.8	58.0–65.3
Posterior dorsal to anterior anal	34.9	33.0	1.4	30.2–35.0
Posterior dorsal to posterior anal	19.9	18.6	1.0	17.0–19.9
Posterior dorsal to ventral caudal	20.9	19.3	0.9	17.5–20.9
Posterior anal to dorsal caudal	23.2	22.0	0.9	21.0–23.8
Anterior dorsal to pelvic-fin origin	42.7	38.5	1.8	35.0–42.7
Posterior dorsal to pelvic-fin origin	61.1	57.5	1.7	54.7–61.1
Caudal peduncle length	13.6	13.3	1.0	11.5–14.8
Least caudal peduncle depth	15.8	15.6	0.7	14.4–16.3
Meristics		Mode	Frequency (%)	Range
Dorsal-fin spines	17	17	80.0	16–18
Dorsal-fin rays	8	8	80.0	7–9
Anal-fin spines	3	3	100.0	3–3
Anal-fin rays	7	8	75.0	7–8
Pectoral-fin rays	14	14	45.0	12–15
Pelvic-fin rays	6	6	90.0	6–7
Lateral line scales	30	30	40.0	29–32
Pored scales caudal	2	1	40.0	0–3
Cheek scale rows	4	4	85.0	4–5
Gill rakers 1st ceratobranchial	10	10	70.0	9–11
Gill rakers 1st epibranchial	5	4	85.0	3–5
Teeth outer left lower jaw	12	10	25.0	6–17
Tooth rows upper jaw	4	3	70.0	3–4
Tooth rows lower jaw	4	3	90.0.	3–4