Two new species of stone loaches of the genus Barbatula (Cypriniformes: Nemacheilidae) from Europe with a neotype designation of B. barbatula (Teleostei: Nemacheilidae) Author Calegari, Bárbara B. Aquatic Ecology and Evolution, Institute of Ecology and Evolution, University of Bern, Bern, Switzerland & Department of Fish Ecology and Evolution, EAWAG, Swiss Federal Institute for Aquatic Science and Technology, Kastanienbaum, Switzerland & Wyss Academy for Nature at the University of Bern, Bern, Switzerland barbara.calegari@unibe.ch Author Freyhof, Jörg Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Berlin, Germany Author Waldock, Conor Aquatic Ecology and Evolution, Institute of Ecology and Evolution, University of Bern, Bern, Switzerland & Department of Fish Ecology and Evolution, EAWAG, Swiss Federal Institute for Aquatic Science and Technology, Kastanienbaum, Switzerland & Wyss Academy for Nature at the University of Bern, Bern, Switzerland Author Wegscheider, Bernhard Aquatic Ecology and Evolution, Institute of Ecology and Evolution, University of Bern, Bern, Switzerland & Department of Fish Ecology and Evolution, EAWAG, Swiss Federal Institute for Aquatic Science and Technology, Kastanienbaum, Switzerland & Wyss Academy for Nature at the University of Bern, Bern, Switzerland Author Josi, Dario Aquatic Ecology and Evolution, Institute of Ecology and Evolution, University of Bern, Bern, Switzerland & Department of Fish Ecology and Evolution, EAWAG, Swiss Federal Institute for Aquatic Science and Technology, Kastanienbaum, Switzerland & Wyss Academy for Nature at the University of Bern, Bern, Switzerland Author Rüber, Lukas Aquatic Ecology and Evolution, Institute of Ecology and Evolution, University of Bern, Bern, Switzerland & Naturhistorisches Museum Bern, Bern, Switzerland Author Seehausen, Ole Aquatic Ecology and Evolution, Institute of Ecology and Evolution, University of Bern, Bern, Switzerland & Department of Fish Ecology and Evolution, EAWAG, Swiss Federal Institute for Aquatic Science and Technology, Kastanienbaum, Switzerland text Journal of Fish Biology 2025 2025-07-02 107 4 1364 1397 journal article 10.1111/jfb.70108 272faa11-1c7b-4955-8172-7f1e74f6b939 1095-8649 17584041

TABLE 1 Morphometric data for Barbatula barbatula, n = 3 specimens, range includes neotype (Neo).

	Neo	Min	Max	Mean	SD
Abbreviations: Max, maximum; Min, minimum; SD, standard deviation.
Standard length (mm)	46.2	39.8	46.2	43.5	-
Percentage of standard length
Dorsal head length	25.8	23.2	25.8	24.3	1.3
Lateral head length	29.6	25.9	29.6	27.8	1.9
Predorsal length	60.4	55.8	60.4	57.5	2.5
Postdorsal length	32.3	32.3	32.4	32.3	0.1
Prepectoral length	28.7	26.2	28.7	27.5	1.3
Prepelvic length	59.5	55.9	59.6	58.3	2.1
Preanal length	83.0	81.0	83.0	82.1	1.0
Pectoral-fin origin to pelvic-fin origin	33.8	30.9	33.8	32.4	1.5
Pelvic-fin origin to anal-fin origin	26.0	23.3	26.0	24.8	1.4
First dorsal-fin ray length	20.0	20.0	24.3	22.4	2.2
Dorsal fin length	25.4	24.1	25.8	25.1	0.9
Dorsal-fin base length	18.6	14.5	18.6	16.0	2.2
Anal-fin base length	11.8	8.3	11.8	9.7	1.8
First pectoral-fin ray length	22.3	17.7	22.3	20.0	3.3
Pectoral fin length	26.7	20.4	26.7	23.5	4.4
First pelvic-fin ray length	14.5	13.1	17.6	15.1	2.3
Pelvic fin length	19.3	17.2	19.3	18.3	1.1
Caudal-peduncle length	12.5	11.7	12.7	12.3	0.5
Caudal-peduncle depth	14.9	9.8	14.9	12.3	2.5
Caudal-peduncle width	2.6	1.7	2.6	2.2	0.4
Body depth at pectoral-fin origin	16.4	14.3	16.6	15.8	1.3
Body width at dorsal-fin origin	12.7	11.8	13.0	12.5	0.6
Percentage of head length
Head depth	54.3	51.7	57.3	54.4	2.8
Snout length	52.2	50.8	52.6	51.9	1.0
Postorbital length	50.9	47.0	54.2	50.7	3.6
Orbital diameter	23.0	19.4	24.7	22.4	2.8
Suborbital depth	36.3	33.1	36.3	34.2	1.8
Interorbital distance	31.9	30.1	34.2	32.0	2.1
Anterior internarial distance	16.0	12.8	16.8	15.2	2.1
Posterior internarial distance	27.1	23.0	27.1	25.5	2.2
Upper – lower lips distance	29.7	23.0	29.7	26.9	3.5
Lower lip length	21.2	16.6	21.2	18.4	2.4
Mental lobe length	9.4	9.3	9.9	9.5	0.3
Mouth width	36.4	31.4	36.4	34.6	2.8