Two new species of stone loaches of the genus Barbatula (Cypriniformes: Nemacheilidae) from Europe with a neotype designation of B. barbatula (Teleostei: Nemacheilidae) Author Calegari, Bárbara B. Aquatic Ecology and Evolution, Institute of Ecology and Evolution, University of Bern, Bern, Switzerland & Department of Fish Ecology and Evolution, EAWAG, Swiss Federal Institute for Aquatic Science and Technology, Kastanienbaum, Switzerland & Wyss Academy for Nature at the University of Bern, Bern, Switzerland barbara.calegari@unibe.ch Author Freyhof, Jörg Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Berlin, Germany Author Waldock, Conor Aquatic Ecology and Evolution, Institute of Ecology and Evolution, University of Bern, Bern, Switzerland & Department of Fish Ecology and Evolution, EAWAG, Swiss Federal Institute for Aquatic Science and Technology, Kastanienbaum, Switzerland & Wyss Academy for Nature at the University of Bern, Bern, Switzerland Author Wegscheider, Bernhard Aquatic Ecology and Evolution, Institute of Ecology and Evolution, University of Bern, Bern, Switzerland & Department of Fish Ecology and Evolution, EAWAG, Swiss Federal Institute for Aquatic Science and Technology, Kastanienbaum, Switzerland & Wyss Academy for Nature at the University of Bern, Bern, Switzerland Author Josi, Dario Aquatic Ecology and Evolution, Institute of Ecology and Evolution, University of Bern, Bern, Switzerland & Department of Fish Ecology and Evolution, EAWAG, Swiss Federal Institute for Aquatic Science and Technology, Kastanienbaum, Switzerland & Wyss Academy for Nature at the University of Bern, Bern, Switzerland Author Rüber, Lukas Aquatic Ecology and Evolution, Institute of Ecology and Evolution, University of Bern, Bern, Switzerland & Naturhistorisches Museum Bern, Bern, Switzerland Author Seehausen, Ole Aquatic Ecology and Evolution, Institute of Ecology and Evolution, University of Bern, Bern, Switzerland & Department of Fish Ecology and Evolution, EAWAG, Swiss Federal Institute for Aquatic Science and Technology, Kastanienbaum, Switzerland text Journal of Fish Biology 2025 2025-07-02 107 4 1364 1397 journal article 10.1111/jfb.70108 272faa11-1c7b-4955-8172-7f1e74f6b939 1095-8649 17584041

TABLE 2 Morphometric data for holotypes (Hol) and paratypes of Barbatula fluvicola (n = 25) and B. ommata (n = 25), range includes holotype.

	B. fluvicola					B. ommata
Abbreviations: Max, maximum; Min, minimum; SD, standard deviation.
	Hol	Min	Max	Mean	SD	Hol	Min	Max	Mean	SD
Standard length (mm)	83.8	64.3	91.2	77.7	-	74.1	55.2	82.0	67.1	-
Percentage of standard length
Dorsal head length	18.9	18.4	21.7	19.6	0.7	20.6	19.3	21.4	20.3	0.6
Lateral head length	24.3	21.8	24.5	23.3	0.7	23.5	22.4	26.2	23.8	0.9
Predorsal length	52.7	50.5	55.8	53.2	1.3	53.3	50.4	56.8	53.6	1.4
Postdorsal length	35.4	34.3	38.0	36.3	1.1	35.2	31.4	38.3	35.6	1.9
Prepectoral length	23.6	20.9	25.1	22.8	0.8	22.4	21.3	25.8	22.9	0.9
Prepelvic length	53.7	51.2	55.9	53.8	1.2	55.4	53.4	58.7	55.5	1.5
Preanal length	79.0	75.4	81.1	77.8	1.3	77.6	76.2	83.8	78.8	1.8
Pectoral-fin origin to pelvic-fin origin	31.3	29.3	34.8	32.2	1.3	34.2	29.7	37.3	33.5	1.6
Pelvic-fin origin to anal-fin origin	25.6	22.2	27.4	24.4	1.5	22.9	21.0	29.5	23.7	1.8
First dorsal-fin ray length	15.4	15.0	21.2	16.9	1.4	13.6	13.5	19.8	16.6	1.8
Dorsal fin length	19.8	18.4	24.2	20.8	1.3	19.3	18.4	23.3	20.8	1.3
Dorsal-fin base length	13.0	11.2	13.6	12.3	0.7	12.9	11.1	14.3	12.6	0.9
Anal-fin base length	8.9	7.2	9.9	8.3	0.7	8.0	7.4	9.4	8.3	0.5
First pectoral-fin ray length	14.1	12.5	21.3	15.2	2.2	13.4	11.9	21.1	15.8	2.3
Pectoral fin length	17.5	17.1	23.4	18.9	1.6	16.8	15.5	23.5	19.1	2.5
First pelvic-fin ray length	9.9	8.5	15.4	11.5	1.5	9.8	9.6	14.5	12.0	1.4
Pelvic fin length	13.4	13.4	18.8	15.4	1.1	14.4	9.9	17.1	15.1	1.5
Caudal-peduncle length	15.2	13.9	17.3	15.8	0.9	12.7	12.7	16.2	14.5	1.1
Caudal-peduncle depth	10.3	8.7	11.8	10.1	0.8	9.9	7.7	10.9	9.1	0.8
Caudal-peduncle width	3.7	2.9	4.9	3.8	0.4	3.0	2.4	4.6	3.3	0.5
Body depth at pectoral-fin origin	13.6	11.4	14.1	12.8	0.7	13.5	11.5	15.0	12.7	0.9
Body width at dorsal-fin origin	12.6	10.5	13.5	12.0	0.7	13.7	9.5	14.1	11.6	1.3
Percentage of head length
Head depth	47.5	46.1	51.7	48.4	1.6	45.3	38.4	46.0	43.5	2.1
Snout length	43.7	39.0	47.6	44.4	1.9	43.3	37.2	46.1	42.0	1.9
Postorbital length	44.5	41.8	48.2	44.8	1.6	41.4	41.4	49.2	43.9	1.6
Orbital diameter	17.5	16.1	19.8	18.3	0.8	23.1	21.1	28.3	23.2	1.6
Suborbital depth	32.1	26.5	32.4	30.1	1.9	29.9	23.7	31.2	27.6	2.0
Interorbital distance	29.1	26.3	32.2	28.5	1.8	31.2	24.0	31.7	29.0	2.0
Anterior internarial distance	13.7	11.8	15.7	13.7	1.1	14.1	10.7	16.3	13.8	1.4
Posterior internarial distance	23.3	21.3	26.5	23.6	1.5	22.9	20.1	30.5	22.8	1.9
Upper – lower lips distance	22.2	20.3	27.3	23.3	1.9	24.8	20.3	26.9	23.7	1.7
Lower lip length	14.9	13.7	17.2	15.9	0.9	15.0	12.7	17.1	15.2	1.0
Mental lobe length	9.9	7.6	12.7	10.3	1.1	8.7	7.3	10.6	9.1	0.8
Mouth width	34.5	31.6	38.3	34.7	1.8	35.6	28.9	36.1	32.8	2.0