Bark beetlesand pinhole borers recently ornewly introduced toFrance (Coleoptera Curculionidae, Scolytinae and Platypodinae) Author Barnouin, Thomas Author Soldati, Fabien 0000-0001-9697-3787 https: // orcid. org / 0000 - 0001 - 9697 - 3787 Author Roques, Alain URZF- Zoologie Forestière, INRAE, 2163 Avenue de la Pomme de Pin, 45075, Orléans, France. Author Faccoli, Massimo Department of Agronomy, Food, Natural Resources, Animals and Environment (DAFNAE), University of Padua, Viale dell’Università, 16, 35020 Legnaro, Italy. Author Kirkendall, Lawrence R. Department of Biology, University of Bergen, P. O. Box 7803, N- 5006 Bergen, Norway. Author Mouttet, Raphaëlle ANSES, Laboratoire de la Santé des Végétaux, 755 avenue du Campus Agropolis, CS 30016, 34988 Montferrier-sur-Lez cedex, France. Author Daubree, Jean-Baptiste 0000-0002-5383-3984 Pôle Sud-Est de la Santé des Forêts, DRAAF SRAL PACA, BP 95, 84141 Montfavet cedex, France. jean-baptiste. daubree @ agriculture. gouv. fr; https: // orcid. org / 0000 - 0002 - 5383 - 3984 jean-baptiste.daubree@agriculture.gouv.fr Author Noblecourt, Thierry 0000-0002-9248-9012 https: // orcid. org / 0000 - 0002 - 9248 - 9012 text Zootaxa 2020 2020-11-09 4877 1 51 74 journal article 8006 10.11646/zootaxa.4877.1.2 02af60c1-b666-4b96-8677-70d2ebc411a4 1175-5326 4423749 3CABEE0D-D1D2-4150-983C-8F8FE2438953

TABLE 2. List of Scolytinae and Platypodinae introduced to France. Phase: see Table 1; Bioclimat: M, Mediterranean region; T, Temperate region. Feeding guild: Spm, spermatophagous, breeding in seeds; Phl, phloeophagous, breeding in inner bark; Xm, xylomycetophagous (ambrosia beetles).

	Native to	Phase	First record	Bioclimat / Feeding guild	Additional informations	References
SCOLYTINAE
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Amasa sp. near truncata (Erichson)	Australia	2	2018	M / Xm	French Riviera:Antibes and Cannes. Host presumably Eucalyptus.	This work
Coccotrypes dactyliperda (Fabricius)	Unknown (cosmopolitan)	5	<1949	M / Spm	Common. Entire Mediterranean region. In seeds of palm trees.	Balachowsky 1949; Kirkendall & Faccoli 2010
Cyclorhipidion bodoanum (Reitter)	North Asia	5	1960	T / Xm	Common. Widespread in the French mainland, especially in plains. On broadleaved trees (mainly oaks).	Kirkendall & Faccoli 2010
Cyclorhipidion distinguendum (Eggers)	Southeast Asia	3	2013	T / Xm	Rare. Alps: Dauphiné. On broadleaved trees (Fagaceae).	Dodelin 2018; this work
Cryphalus dilutus Eichhoff	Indian subcontinent	2	2017	M/ Phl	French Riviera: Saint-Tropez area. On broadleaved trees (fig trees).	This work
Dactylotrypes longicollis (Wollaston)	Canary Islands	5	1940	M / Spm	Locally common invasive throughout Mediterranean. In the seeds of palm trees.	Balachowsky 1949; Kirkendall & Faccoli 2010
Dryocoetes himalayensis Strohmeyer	South Asia (India)	4	1975	T / Phl	Rare. Sporadic in France. On broadleaved trees (walnut and probably oaks).	Knižek, 2011a; Schott 2017; this work
Gnathotrichus materiarius (Fitch)	Northeastern USA	5	1933	T / Xm	Common. Widespread in the French mainland, especially plains. On Conifers (mainly pines).	Balachowsky, 1949; Kirkendall & Faccoli 2010
Hypothenemus eruditus Westwood	Unknown (cosmopolitan)	5	<1949	M / Phl	Southern France. On broadleaved trees (mainly fig trees).	Balachowsky 1949; Kirkendall & Faccoli 2010
Liparthrum mandibulare (Wollaston)	Canary Islands	2	2015	T / Phl	Locally abundant. French Britany (Côte d’Armor). On various broadleaved trees and shrubs.	Lagarde & Noblecourt 2018
Phloeosinus rudis Blandford	East Asia	1	1940	Phl	Rare. Southeast France: Var. Not recorded since 1945. On Conifers (Cupressaceae).	Balachowsky 1949; Kirkendall & Faccoli 2010
Phloeotribus liminaris (Harris)	East USA	1	2007	Phl	Southwest France: Gironde. On Prunus serotina.	Noblecourt & Lessieur 2016
Xyleborus affinis Eichhoff	Unknown (cosmotropical)	1	2016	Xm	Charente-Maritime: port of La Rochelle.	This work
Xyleborus bispinatus Eichhoff	American tropics?	2	2017	M / Xm	French Riviera: Nice. Broadleaved trees.	This work
Xyleborus ferrugineus (Fabricius)	American tropics?	1	2016	Xm	Charente-Maritime: port of La Rochelle. Broadleaved trees.	This work
Xyleborus pfeilii (Ratzeburg)	East Asia	5	19th century	T / Xm	Very rare. Here and there in France. On broadleaved trees.	Balachowsky 1949; Kirkendall & Faccoli 2010
Xylosandrus compactus (Eichhoff)	Southeast Asia	4	2015	M / Xm	Locally abundant. Southeast France: Alpes- Maritimes and Var. On broadleaved trees.	Chapin et al. 2016; this work
Xylosandrus crassiusculus (Motschulsky)	Asian tropics	4	2013	M / Xm	Locally abundant. South of France. On broadleaved trees.	This work
Xylosandrus germanus (Blandford)	East Asia	5	1984	T (M) / Xm	Very common. Widespread in nearly all France. On broadleaved trees.	Schott 1994; Nageleisen et al. 2015; Kirkendall & Faccoli 2010
Xylosandrus morigerus (Blandford)	Asian tropics?	1	1897	Xm	Found in greenhouses in Marseille. Not recorded since 19th century. On orchids (Dendrobium phalaenopsis).	Balachowsky 1949; Kirkendall & Faccoli 2010
Xyloterinus politus (Say)	North America	2	2017	T / Xm	Rare. Northwestern France: between Le Havre and Rouen (Seine-Maritime). On broadleaved trees (field maple).	Dodelin & Saurat 2017; this work
PLATYPODINAE
Euplatypus hintzi (Schaufuss )	African tropics	1	2012	Xm	Charente-Maritime: port of La Rochelle, Herault: port of Sète. Polyphagous.	This work
Euplatypus parallelus (Fabricius)	American tropics	1	2012	Xm	Charente-Maritime: port of La Rochelle. Polyphagous.	This work