Systematics of the Calotes irawadi complex (Squamata, Agamidae) with two newly described species from Thailand Author Prakobkarn, Arpapan https://orcid.org/0009-0000-9047-1102 Ph. D. Program in Zoology, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand Author Zug, George R. 0000-0003-4922-5721 Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20560, USA Author Tandavanitj, Nontivich Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand Author Ngamprasertwong, Thongchai 0000-0003-3146-1849 Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand text ZooKeys 2026 2026-06-03 1281 69 104 journal article 315339 10.3897/zookeys.1281.175455 1321a16e-cff9-4247-8200-6557d35c47d3 E54EA084-1419-444B-B507-C4646540A63B

Table 5. Morphological (meristic) characters of holotype, paratypes and all specimens of Calotes thailandensis sp. nov., C. maehongsonensis sp. nov., C. irawadi, and C. wangi. Numbers of scales are represented in mean ± SD (range). Abbreviations are showed in Materials and methods.

Characters	C. thailandensis sp. nov.					C. maehongsonensis sp. nov.					C. irawadi		C. wangi
	Holotype	Paratypes		All specimens		Holotype	Paratypes		All specimens		All specimens		Huang et al. (2023)
	Male	Male (n = 5)	Female (n = 5)	Male (n = 26)	Female (n = 27)	Male	Male (n = 5)	Female (n = 5)	Male (n = 10)	Female (n = 10)	Male (n = 28)	Female (n = 14)	Male (n = 2)
* unknown sample size.
CanthR	8	7.8 ± 0.4	7.6 ± 0.6	8.0 ± 0.5	7.6 ± 0.5	8	8.2 ± 0.4	8 ± 0.0	8.2 ± 0.4	8.0 ± 0	7.9 ± 0.6	7.7 ± 0.5	4.91 *
		(7–8)	(7–8)	(7–9)	(7–8)		(8–9)	8	(8–9)	8	(6–9)	(7–8)	(4–5)
Eyelid	11	12.2 ± 1.3	9.8 ± 0.8	12.2 ± 1.1	9.6 ± 1.1	14	12.4 ± 0.5	9.2 ± 1.3	12.4 ± 1.1	9.5 ± 1.1	12.7 ± 1.4	11.9 ± 0.7	12.4 *
		(11–14)	(9–11)	(10–14)	(8–12)		(12–13)	(8–11)	(11–14)	(8–11)	(10–15)	(11–13)	(10–14)
HeadSLn	13	13.4 ± 1.7	12.6 ± 0.6	13.4 ± 1.0	13.2 ± 1.3	14	14.4 ± 0.5	12.8 ± 1.8	14.3 ± 0.5	13.4 ± 1.6	12.1 ± 1.2	14.0 ± 1.5
		(12–16)	(12–13)	(12–16)	(11–17)		(14–15)	(11–15)	(14–15)	(11–16)	(10–14)	(12–17)
HeadSTr	15	13.0 ± 0.7	13.4 ± 1.5	13.8 ± 1.0	13.0 ± 1.3	13	13.0 ± 0.7	12 ± 0.7	13.1 ± 0.7	12.8 ± 2.3	13.9 ± 1.2	12.0 ± 1.0
		(12–14)	(11–15)	(12–15)	(11–16)		(12–14)	(11–13)	(12–14)	(11–19)	(12–16)	(10–13)
Inflab	10	10.0 ± 0.0	9.6 ± 0.9	10.1 ± 0.7	9.3 ± 0.6	10	10.4 ± 0.5	9.4 ± 0.9	10.1 ± 0.7	9.2 ± 0.9 b	10.3 ± 0.7	10.1 ± 1.1
		10	(9–11)	(9–11)	(8–11)		(10–11)	(8–10)	(9–11)	(8–10)	(9–12)	(9–12)	(9–12)
Suplab	10	10.4 ± 0.5	9.2 ± 0.5	10.5 ± 0.6	9.1 ± 0.5	11	11.0 ± 0.0	9.8 ± 0.8	10.8 ± 0.4	9.7 ± 0.7	10.9 ± 0.7	10.6 ± 0.9
		(10–11)	(9–10)	(9–12)	(8–10)		11	(9–11)	(10–11)	(9–11)	(9–12)	(9–13)	(9–11)
SnS	7	6.4 ± 0.5	6.2 ± 0.5	6.76 ± 0.7	6.4 ± 0.5	5	6.0 ± 0.7	6.2 ± 0.4	6.0 ± 0.7	6.1 ± 0.3	6.8 ± 0.6	6.9 ± 0.8
		(6–7)	(6–7)	(6–8)	(6–7)		(5–7)	(6–7)	(5–7)	(6–7)	(6–8)	(6–8)
4 FingLm	21	19.8 ± 1.1	19.6 ± 1.1	20.1 ± 1.2	19.4 ± 1.0	19	19.8 ± 0.8	19 ± 1.6	19.5 ± 0.8	18.6 ± 1.3	20.9 ± 1.4	19.4 ± 1.4
		(19–21)	(18–21)	(18–22)	(17–21)		(19–21)	(17–21)	(18–21)	(17–21)	(18–24)	(17–22)	(20–22)
4 ToeLm	23	25.2 ± 2.2	25.0 ± 1.0	25.1 ± 1.7	24.8 ± 1.3	23	24.6 ± 1.1	23.4 ± 1.5	24.2 ± 1.1	22.5 ± 1.5 b	25.3 ± 1.6	24.1 ± 1.4	24.2 *
		(23–28)	(24–26)	(23–29)	(23–28)		(23–26)	(21–25)	(23–26)	(21–25)	(23–29)	(22–28)	(19–27)
Midbody	43	41.6 ± 2.7	42.4 ± 1.8	42.5 ± 2.0	44.3 ± 3.0	44	44.6 ± 0.5	44.2 ± 2.5	44.5 ± 0.8	43.5 ± 2.0	45.4 ± 2.1	46.4 ± 2.3	41.2 *
		(37–44)	(40–44)	(37–46)	(39–51)		(44–45)	(42–48)	(43–46)	(41–48)	(42–50)	(43–51)	(36–46)
Dorsal	46	46.6 ± 1.8	46.6 ± 1.7	46.0 ± 1.9	47.2 ± 1.8	51	48.2 ± 1.8	51 ± 2.4	48.8 ± 1.5	50.8 ± 2.1 a	48.3 ± 2.4	51.6 ± 4.5	12.4
		(45–49)	(45–49)	(40–49)	(44–50)		(47–51)	(48–54)	(47–51)	(48–54)	(41–53)	(45–59)	(38–44)