Two new species of Cyrtodactylus (Squamata: Gekkonidae) from northern Laos including new finding and expanded diagnosis of C. bansocensis Author Schneider, Nicole 0000-0002-6588-229X AG Zoologischer Garten Köln, Riehler Strasse 173, D- 50735 Cologne, Germany. https: // orcid. org / 0000 - 0002 - 4797 - 609 X. & Institute of Zoology, University of Cologne, Zülpicher Strasse 47 b, D- 50674 Cologne, Germany. schneider. nicole 87 @ gmail. com; https: // orcid. org / 0000 - 0002 - 6588 - 229 X. schneider.nicole87@gmail.com Author Luu, Vinh Quang 0000-0002-0634-1338 Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Xuan Mai, Chuong My, Hanoi, Vietnam. qvinhfuv @ yahoo. com. au; https: // orcid. org / 0000 - 0002 - 0634 - 1338. qvinhfuv@yahoo.com.au Author Sitthivong, Saly 0000-0002-9993-2136 Faculty of Forest Resources and Environmental Management, Vietnam National University of Forestry, Xuan Mai, Chuong My, Hanoi, Vietnam. qvinhfuv @ yahoo. com. au; https: // orcid. org / 0000 - 0002 - 0634 - 1338. & Faculty of Forestry, National University of Laos, Dong Dok Campus, Vientiane, Lao PDR. sitthivong _ frt @ yahoo. co. th; https: // orcid. org / 0000 - 0002 - 9993 - 2136 qvinhfuv@yahoo.com.au Author Teynié, Alexandre 0000-0002-6588-229X Société d’Histoire Naturelle Alcide d’Orbigny, 57, rue de Gergovie, 63170 Aubière, France. ateynie @ shnao. net; https: // orcid. org / 0000 - 0002 - 6588 - 229 X. ateynie@shnao.net Author Le, Minh Duc 0000-0002-2953-2815 Faculty of Environmental Sciences, University of Science, Vietnam National University Hanoi, 334 Nguyen Trai Road, Hanoi, Vietnam. le. duc. minh @ hus. edu. vn; https: // orcid. org / 0000 - 0002 - 2953 - 2815. & Central Institute for Natural Resources and Environmental Studies, Vietnam National University Hanoi, 19 Le Thanh Tong, Hanoi, Vietnam. & Department of Herpetology, American Museum of Natural History, Central Park West at 79 Street, New York, New York 10024 Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam. nqt 2 @ yahoo. com; https: // orcid. org / 0000 - 0002 - 6601 - 0880 le.duc.minh@hus.edu.vn Author Nguyen, Truong Quang 0000-0002-6601-0880 nqt2@yahoo.com Author Ziegler, Thomas 0000-0002-4797-609X AG Zoologischer Garten Köln, Riehler Strasse 173, D- 50735 Cologne, Germany. https: // orcid. org / 0000 - 0002 - 4797 - 609 X. & Institute of Zoology, University of Cologne, Zülpicher Strasse 47 b, D- 50674 Cologne, Germany. schneider. nicole 87 @ gmail. com; https: // orcid. org / 0000 - 0002 - 6588 - 229 X. schneider.nicole87@gmail.com text Zootaxa 2020 2020-08-07 4822 4 503 530 journal article 8764 10.11646/zootaxa.4822.4.3 67b665b5-74e4-4d65-b375-a07f33a7aa75 1175-5326 4452362 4C7F87B9-D1C1-4821-AB37-BBA3FA59F55C

TABLE 2. Morphological characters, distribution and habitat information of all Laotian Cyrtodactylus species. Abbreviations are as follows: SVL: snout–vent length (in mm); TaL: Tail length (in mm); SPL: Supralabials; IFL: Infralabials; SMC: along the midbody from mental shield to anterior edge of cloaca; MS: midbody scales; V: ventral scales; DTR: dorsal tubercle rows; EPS: enlarged precloacal scales; EFS: enlarged femoral scales; FP: femoral pores; PP: precloacal pores; LD4/LT4: subdigital lamellae under fourth finger/toe; *=regenerated tail;?=unknown character.

Taxa	max. SVL	max. TaL	SPL	IFL	SMC	MS	V	DTR	EFS	LD4	LT4	Color pattern of dorsum	Enlarged subcaudals	FP	PP (in males)	PP (in females)	Distribution (Laos)	Habitat
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Cyrtodactylus houaphanensis sp. nov.	75.8	59.1*	9–10	8–9	184	127	35	20	absent	16–18	19–23	banded	present	absent	6	?	Houaphan	karst forest
Cyrtodactylus ngoiensis sp. nov.	95.3	101.8	6–9	8–11	179–199	102–121	38–43	15–21	present	16–18	19–20	banded	present	14 in male/ 0 in females	7	7 (pitted)	Luang Prabang	karst forest
C. bansocensis	74.0	103.5	8–12	8–10	158–204	86–102	34–35	14–20	present	16–20	18–21	banded	present	present	34 (PP+FP)	16 (PP+FP)	Khammouane	karst forest
C. buchardi	65.0	54.0	13–14	10–11	?	69	30	25	absent	14	12	blotched	absent	absent	9	0	Champasak	lowland monsoon evergreen forest
C. calamei	89.3	107.5	9–11	8–11	183–193	101–114	39–42	10–16	present	17	19/21	banded	present	present	35–39 (PP+FP)	38 (PP+FP)	Khammouane	karst forest
C. cryptus	83.6	73.0	7–8	6–7	211–218	122–131	40–43	15–17	absent	17–19	17–19	blotched	absent	absent	8–11	8 (pitted)	Khammouane	karst forest
C. darevskii	100.0	113.0	10–12	9–11	180–216	?	38–46	16–20	present	17–20	18–22	banded	present	present	38–44 (PP+FP)	24–34 (PP+FP)	Khammouane	karst forest
C. hinnamnoensis	100.6	108.3	9–12	7–11	179–201	93–112	35–48	14–19	present	16–21	18–22	banded	present	present	36–44 (PP+FP)	0–28 (PP+FP)	Khammouane	karst forest
C. interdigitalis	80.0	95.2	11–15	9–12	196	112	37–42	18–22	present	16–22	16–20	banded	present	17 in males/ 0 in females	14	0–15 (pitted)	Vientiane, Khammouane	karst forest/ primary forest
C. jaegeri	68.5	83.4	10–11	9–11	156–164	?	31–32	15–17	17–19	17–19	20–23	banded	present	present	44 (PP+FP)	21 (FP+PP pitted)	Khammouane	karst forest
C. jarujini	90.0	116.0	11–16	10–12	169	?	32–38	16–17	present	15–18	18–20	blotched	present	present in males	52–57 (PP+FP)	0	Bolikhamxay	lowland monsoon evergreen forest
C. khammouanensis	73.0	95.0	11–12	9–10	155–172	?	32–38	16–21	present	18–20	20–23	banded	present	present in males	40–44 (PP+FP)	0–17	Khammouane	karst forest
C. lomyenensis	71.2	86.9	10–14	9–11	190–205	84–93	35–40	19–24	17–18	16–21	18–24	banded	present	present	39–40 (PP+FP)	32 (PP+FP)	Khammouane	karst forest
C. muangfuangensis	83.9	102.6	9–11	9–11	176–182	102–112	31–37	15–16	present	16–19	18–23	banded	present	present	8+6+7 (FPl+PP+FPr)	5–9+6+5–8 (FPl+PP+FPr (pitted scales)	Vientiane	karst forest
C. multiporus	98.0	115.2	9–11	8–11	164–182	?	30–38	15–20	absent	16–20	18–22	blotched	present	present in males only	50–60 (PP+FP)	0–14 (pitted)	Khammouane	karst forest
C. pageli	87.7	113.2	9–12	9–10	216–239	113–124	41–46	9–14	absent	19–23	19–26	banded	present	absent	4	4–6	Vientiane	karst forest
C. pseudoquadrivirgatus	83.8	82.5	8–13	7–10	?	?	39–57	16–24	absent	15–21	16–25	blotched	absent	absent	5–9	5–9	Salavan	evergreen forest
C. rufford	72.5	96.8	11–12	9–11	153–167	74–79	27–29	14–16	present	19–20	18–19	banded	present	present	42–43 (PP+FP)	?	Khammouane	karst forest
C. sommerladi	80.3	89.9	9–11	8–9	168–192	76–93	31–39	0–5	present	16–20	17–24	banded	present	present	20–26 (PP+FP)	17–21	Khammouane	karst forest
C. soudthichaki	70.0	95.2	10–11	8–9	165–170	78–85	32–33	19–20	present	16–18	18	banded	present	present	29 (PP+FP)	0	Khammouane	karst forest
C. spelaeus	91.0	83.0	9–12	8–10	156–183	?	36–39	10	absent	19–20	22–24	blotched	present	absent	8–9	0	Vientiane	karst forest
C. thathomensis	75.5	96.0	10–11	9–10	?	?	30–36	14–18	present	16–17	18–20	banded	present	present	10	10–12 (pitted)	Xiangkhoang	Karst forest
C. teyniei	89.9	110.0	10	9	184	108	38	18–22	present	17–18	19–20	blotched	present	Absent in females, unknown in males	unknown	14	Bolikhamxay	karst forest
C. vilaphongi	86.1	68.1	9–10	7–9	161–165	106–122	34–36	15–16	absent	18–19	18–20	banded	absent	?	?	0	Luang Prabang	karst forest
C. wayakonei	86.8	89.0	7–8	9–10	151–163	85–98	31–35	17–19	absent	17–18	19–20	banded	absent	absent	6–8	7	Luang Nam Tha	karst forest